Specimen of Nesopupa, photographed by John Slapcinsky.

Belongs within: Pupilloidea.

The Nesopupinae are a group of cylindric small snails in which the penis bears an appendix and the retractor muscle is forked. The genus Nesopupa, found in the Indo-Pacific region, has a golden-brown shell with a granular to wrinkled sculpture, and varying numbers of parietal and palatal teeth in the aperture (Stanisic et al. 2010).

<==Nesopupinae [Nesopupidae] BR05
    |--Cylindrovertilla Boettger 1881 BR05, BR17 [incl. Wallivertilla Iredale 1937 SS10; Cylindrovertillidae]
    |    |--*C. fabreana (Crosse 1872) [=Pupa fabreana] SS10
    |    |--C. hedleyi Pilsbry 1920 [incl. C. fabreana boynensis Iredale 1937] SS10
    |    `--C. kingi (Cox 1864) [incl. Pupa mastersi Cox 1864, Wallivertilla kingi negata Iredale 1940] SS10
    `--Nesopupa Pilsbry 1900 [incl. Somniopupa Iredale 1937, Westralcopta Iredale 1939; Nesopupini] SS10
         |--*N. tantilla (Gould 1847) [=Pupa tantilla] SS10
         |--N. barrackporensis (Gude 1914) S88
         |--N. dentifera (Pease 1871) S88
         |--N. ganzae Adam 1954 S88
         |--N. hackerae Stanisic in Stanisic, Shea et al. 2010 SS10
         |--N. insularis (Darteville 1952) S88
         |--N. madgei Peile 1936 S88
         |--N. moluccana (Boettger 1891) S88
         |--N. moreleti (Brown 1870) S88
         |--N. novopommerana Rensch 1932 SS10 [incl. N. tenimberica Haas 1937 SS10, N. (Insulipupa) tenimberica S88]
         |--N. peilei Madge 1938 S88
         |--N. quadrasi (Moellendorff 1894) S88
         |--N. rodriguezensis Connolly 1925 S88
         |--N. scotti (Brazier 1875) SS10 (see below for synonymy)
         `--N. selebensis (Tapparone-Canefri 1884) S88

Nesopupa scotti (Brazier 1875) SS10 [=Somniopupa scotti S88; incl. Pupa mooreana Smith 1894 SS10, Australbinula (*Westralcopta) mooreana S88, Bifidaria mooreana S88, Gastrocopta mooreana S88, Nesopupa mooreana S88]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

Last updated: 20 January 2020.

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