Helicina concentrica, from here.

Belongs within: Neritimorpha.

Helicinoidea is a clade recognised by molecular analysis containing the terrestrial Helicinidae and the Neritiliidae, found in fresh water and marine caves. No definite morphological synapomorphies have been identified, though both families have the vaginal and ootype openings widely separated, and the vaginal opening situated deep inside the pallial cavity (Kano et al. 2002).

<==Helicinoidea [Helicinacea] BR05
    |--Dawsonella Bradley 1874 [Dawsonellidae, Dawsonellinae] BR05
    |--Deianira Stoliczka 1860 [=Dejanira (l. c.); Deianiridae, Dejaniridae] BR05
    |--Proserpinellidae BR05
    |    |--Proserpinella Bland 1865 BR05
    |    `--Ceres Gray 1856 [Cererinae, Ceresidae, Ceresinae] BR05
    |--Proserpinidae [Proserpininae] BR05
    |    |--Despoena Newton 1891 [Despoenidae] BR05
    |    `--Proserpina Sowerby 1839 BR05
    |--Neritiliidae [Neritiliinae] BR05
    |    |--Pisulinella Kano & Kase 2000 B01
    |    |    `--*P. miocenica B01
    |    |--Pisulina Nevill & Nevill 1869 B01
    |    |    |--*P. adamsiana Nevill & Nevill 1869 B01
    |    |    |--P. biplicata Thiele 1925 B01
    |    |    |--P. maxima Kano & Kase 2000 B01
    |    |    `--P. subpacifica B01
    |    `--Neritilia Martens 1879 BR05
    |         |--*N. rubida (Pease 1865) [=Neritina rubida; incl. Neritilia hawaiiensis Kay 1979] B01
    |         |--N. consimilis (Martens 1879) B01
    |         |--N. monoeli Dohrn 1866 B01
    |         |--N. panamensis B01
    |         `--N. succinea (Récluz 1841) B01
    `--Helicinidae BR05
         |  i. s.: Pleuropoma Moellendorff 1893 [incl. Aphanoconia Wagner 1905, Sphaeroconia Wagner 1905] S88
         |           |--P. draytonensis SC07
         |           |--P. jana SC07
         |           |--P. louisiadensis (Forbes 1851) S88
         |           |--P. macleayi (Brazier 1876) S88
         |           |--P. solomonensis (Smith 1885) S88
         |           `--P. walkeri (Smith 1894) KKW12 (see below for synonymy)
         |         Ceratopoma Moellendorff 1893 S88
         |         Geophorus AA88
         |           |--G. bothropoma Moellendorff 1895 AA88
         |           `--G. trochiformis (Sowerby 1842) AA88
         |         Alcadia BC01
         |           |--A. delmonteana Pilsbry 1932 BC01
         |           `--A. dysonia GO06
         |         Lucidella (Poeniella) peasei Pilsbry 1933 BC01
         |         Palaeohelicina (Palaeohelicina) mayri Clench 1958 BC01
         |--Ceratodiscus Simpson & Henderson 1901 [Ceratodiscinae] BR05
         |--Dimorphoptychia Sandberger 1871 [Dimorphoptychiinae] BR05
         |--Hendersonia Wagner 1905 [Hendersoniinae] BR05
         |--Viana Adams & Adams 1856 [Vianinae] BR05
         |--Stoastoma Adams 1849 [Stoastomatidae, Stoastomatinae, Stoastomidae] BR05
         |    `--*S. pisum Adams 1849 BC01
         `--Helicininae BR05
              |--Bourciera Pfeiffer 1852 [Bourcierinae] BR05
              |--Olygyra Say 1818 [=Oligyra; Oligyradae, Olygyridae] BR05
              |    `--O. sprucei [=Helicina (Oligyra) sprucei] A66
              `--Helicina Lamarck 1799 [Helicinides] BR05
                   |--H. chionea Pilsbry 199 BC01
                   |--H. (Pachystoma) concentrica A66
                   |--H. hollandi Adams 1849 BC01
                   |--H. laciniosa Mighels 1845 [=Pleuropoma laciniosa; incl. Orobophana berniceia] J49
                   `--H. rotelloidea Mighels 1845 [incl. Pleuropoma hawaiiensis] J49

Pleuropoma walkeri (Smith 1894) KKW12 [=Helicina walkeri KKW12, Orobophana walkeri S88, Aphanoconia (Sphaeroconia) baudinensis Wagner 1909 KKW12]

*Type species of generic name indicated


[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440-445.

[AA88] Auffenberg, K. & T. Auffenberg. 1988. Density, spatial distribution, activity patterns, and biomass of the land snail, Geophorus bothropoma Moellendorff (Prosobranchia: Helicinidae). Nautilus 102 (1): 40-45.

[B01] Bandel, K. 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen des Geologisch-Paläontologischen Institutes der Universität Hamburg 85: 65-164.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[GO06] Giribet, G., A. Okusu, A. R. Lindgren, S. W. Huff, M. Schrödl & M. K. Nishiguchi. 2006. Evidence for a clade composed of molluscs with serially repeated structures: Monoplacophorans are related to chitons. Proceedings of the National Academy of Sciences of the USA 103 (20): 7723-7728.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213-231.

Kano, Y., S. Chiba & T. Kase. 2002. Major adaptive radiation in neritopsine gastropods estimated from 28S rRNA sequences and fossil records. Proceedings of the Royal Society of London Series B—Biological Sciences 269: 2457-2465.

[KKW12] Köhler, F., V. Kessner & C. Whisson. 2012. New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia. Records of the Western Australian Museum 27 (1): 21-39.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455-604.

[SC07] Stanisic, J., R. A. D. Cameron, B. M. Pokryszko & J. C. Nekola. 2007. Forest snail faunas from S. E. Queensland and N. E. New South Wales (Australia): patterns of local and regional richness and differentiation. Malacologia 49 (2): 445-462.

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