Belongs within: Amphipoda.
Contains: Hadziidae, Melita, Maera.
The Melitoidea are a group of gammaroid amphipods found mostly in marine and brackish water.
Characters (from Bousfield 1977): Body not carinate or rostrate, toothed on abdomen only (rarely on peraeon); urosome dorsal spine groups weak or lacking; sexual dimorphism strongly expressed in body size, gnathopods, antenna II and peraeopods; antennae II lacking calceoli; antennae strongly developed, I usually much longer than II, peduncular segment II elongate; accessory flagellum prominent (occasionally very reduced or lacking); inferior antennal sinus small or sharply incised; eye (when present) small, rounded, occasionally reniform. Mouthparts basic: mandibular palp slender, weakly armed, occasionally absent, segment 3 not greatly shortened; lower lip with inner lobes variously developed, often strong; maxillae with plates often small, setose apically; outer plate of maxilla I with 11-7 apical spine-teeth; maxilliped plates moderately strong, marginally spinose, palp dactylate. Coxal plates I-IV medium deep to shallow, contiguous, weakly setose; coxae V-VII with posterior lobe not deeper than anterior. Gnathopods I and II subchelate, II much the larger and more powerful (especially in male) and of different form (especially in carpus and propodus); peraeopod III larger than IV, not sexually dimorphi; peraeopods V-VII with bases variously expanded, often sublinear, VII usually longest; pleopods usually well developed, peduncles slender; epimeral plates posteriorly acute. Peduncle of uropod I with baso-facial spine strongly developed; uropod III with rami variably developed, spinose, seldom setose; telson variously bilobed, lobes usually divergent, apices acute, spinose in apical notch. Coxal gills simple, often pedunculate, lacking on peraeon VII; sternal gills absent; brood plates linear, often small, margines few- and short-setose.
<==Melitoidea S86
| i. s.: Parapherusa Stebbing 1906 K-SV02, B77
| `--*P. crassipes (Haswell 1879) [=Harmonia crassipes] B77
|--Hadziidae MD01
|--Carangoliopsis Ledoyer 1970 [Carangoliopsidae] B77
| `--*C. spinulosa Ledoyer 1970 B77
|--Metacrangonyx Chevreux 1909 B77, R86 [incl. Afrocrangonyx Karaman 1981 R86; Metacrangonyctidae MD01]
| |--M. longicaudus Ruffo 1954 R86
| |--M. longipes Chevreux 1909 R86
| |--M. panousei Ruffo 1953 R86
| |--M. sinaicus Ruffo 1982 R86
| `--M. spinicaudatus Karaman & Pesce 1980 R86
`--Melitidae S86
| i. s.: Allomelita pellucida (Sars 1882) S86
| Apoweckelia serrata Stock 1985 S86
| Pintaweckelia grandis Stock 1985 S86
| Radaweckelia brevicauda Stock 1985 S86
| Tagua aporema Lowry & Fenwick 1983 S86
| Zhadia subantarctica Lowry & Fenwick 1983 S86
| Zombiweckelia parvipalpus Stock 1985 S86
| Allotexiweckelia Holsinger 1980 S86
| `--A. hirsuta Holsinger 1980 S86
| Anchialella Barnard 1979 S86
| `--A. vulcanella Barnard 1979 S86
| Galapsiellus Barnard 1976 S86
| `--G. lelouporum (Monod 1970) S86
| Holsingerius Barnard & Karaman 1982 S86
| `--H. samacos (Holsinger 1980) S86
| Indoniphargus Straškraba 1967 S86
| `--*I. indicus (Chilton 1923) B77, S86 [=Niphargus indicus B77]
| Madapisella Stock 1980 S86
| `--M. madagascariensis (Ledoyer 1968) S86
| Nippopisella Stock 1980 S86
| `--N. nagatai (Gurjanova 1965) S86
| Phreatomelita Ruffo 1979 S86
| `--P. paceae Ruffo 1979 S86
| Protohadzia Zimmerman & Barnard 1977 S86
| `--P. schoenerae (Fox 1973) S86
| Texiweckelia Holsinger 1980 S86
| `--T. texensis (Holsinger 1973) S86
| Texiweckeliopsis Barnard & Karaman 1982 S86
| `--T. insolita (Holsinger 1980) S86
| Tunisopisa Stock 1980 S86
| `--T. seurati (Gauthier 1936) S86
| Giniphargus Barnard & Karaman 1979 W86
| `--G. pulchellus (Sayce 1900) [=Niphargus pulchellus] W86
| Saliweckelia Stock 1977 S86
| |--S. emarginata Stock 1977 S86
| `--S. holsingeri Stock 1977 S86
| Crangoweckelia S86
| |--C. mixta Stock 1985 S86
| `--C. spinicauda Stock 1985 S86
| Bahadzia S86
| |--B. latipalpus Stock 1985 S86
| |--B. stocki Holsinger 1985 S86
| `--B. williamsi Holsinger 1985 S86
| Victoriopisa Karaman & Barnard 1979 S86
| |--V. atlantica Stock & Platvoet 1981 S86
| |--V. australiensis (Chilton 1923) S86
| |--V. chilkensis (Chilton 1921) S86
| `--V. epistomata (Griffiths 1974) S86
| Metahadzia Stock 1977 S86
| |--M. adriatica Pesce 1979 S86
| |--M. helladis Pesce 1980 S86
| |--M. minuta (Ruffo 1947) S86
| |--M. pachypoda (Ruffo 1982) S86
| `--M. tavaresi (Mateus & Mateus 1972) S86
| Pseudoniphargus Chevreux 1901 S86
| |--*P. africanus (Chevreux 1901) [=Gammarus africanus] B77
| |--P. adriaticus Karaman 1955 S86
| |--P. branchiatus Stock 1980 S86
| |--P. brevipedunculatus Stock 1980 S86
| |--P. elongatus Stock 1980 S86
| |--P. longispinum Stock 1980 S86
| |--P. macrotelsonis Stock 1980 S86
| |--P. mateusorum Stock 1980 S86
| `--P. unisexualis Stock 1980 S86
| Nedsia Barnard & Williams 1995 FJK08
| |--N. douglasi Barnard & Williams 1995 FJK08
| `--N. straskraba Bradbury & Williams 1996 FJK08
| Norcapensis EHH05
|--+--Melitoides Gurjanova 1934 B77
| | `--*M. makarovi Gurjanova 1934 B77
| |--Gammarella Bate 1857 B77
| | `--*G. fucicola (Leach 1814) [=Pherusa fucicola] B77
| |--Netamelita Barnard 1962 B77
| | `--*N. cordata Barnard 1962 B77
| |--Rotomelita Barnard 1977 B77
| | |--*R. ana Barnard 1977 B77
| | `--R. lokoa Barnard 1977 S86
| |--Paraniphargus Tattersall 1925 B77
| | |--*P. ruttneri Schellenberg 1931 B77, S86
| | `--P. annandalei Tattersall 1925 S86
| |--Eriopisa Stebbing 1890 B77
| | |--*E. elongata (Bruzelius 1859) B77
| | |--E. inaequicaudata Ledoyer 1982 S86
| | |--E. incisa McKinney, Kalke & Holland 1978 S86
| | `--E. philippensis (Chilton 1921) S86
| |--Psammogammarus Karaman 1955 B77
| | |--*P. coecus Karaman 1955 B77, S86 [incl. Eriopisa peresi Ledoyer 1968 S86]
| | |--P. caesicolus Stock 1980 S86
| | |--P. garthi (Barnard 1952) S86
| | |--P. gracilis (Ruffo & Schiecke 1976) S86
| | |--P. longiramus (Stock & Nijssen 1965) S86
| | `--P. scopulorum Stock 1983 S86
| |--Eriopisella Chevreux 1920 B77
| | |--*E. pusilla Chevreux 1920 B77
| | |--E. capensis (Barnard 1916) S86
| | |--E. epimera Griffiths 1974 S86
| | |--E. propagatio Clark Imbach 1967 S86
| | |--E. sechellensis (Chevreux 1901) S86
| | `--E. upolu Barnard 1970 S86
| `--Melita B77
`--+--Bathyceradocus Pirlot 1934 B77
| `--*B. stephenseni Pirlot 1934 B77
|--Ceradocus Costa 1853 B77
| |--*C. orchestiipes Costa 1853 B77
| `--C. semiserratus (Bate 1862) BD95
|--Ceradocoides Nicholls 1938 B77
| `--*C. chiltoni Nicholls 1938 B77
|--Ceradocopsis Schellenberg 1926 B77
| `--*C. kergueleni Schellenberg 1926 B77
|--Paraceradocus Stebbing 1899 B77
| `--*P. miersii (Pfeffer 1888) [=Megamoera miersii] B77
|--Metaceradocoides Birstein & Vinogradov 1960 B77
| `--*M. vitjazi Birstein & Vinogradov 1960 B77
|--Maeracunha Stephensen 1949 B77
| `--*M. tristanensis Stephensen 1949 B77
|--Maeropsis Chevreux 1919 B77
| `--*M. perrieri Chevreux 1919 B77
|--Meximaera Barnard 1969 B77
| `--*M. diffidentia Barnard 1969 B77
|--Jerbarnia Croker 1971 B77
| `--*J. mecochira Croker 1971 B77
|--Anelasmopus Oliveira 1953 B77
| `--*A. kraui Oliveira 1953 B77
|--Parelasmopus Stebbing 1888 B77
| `--*P. suluensis (Dana 1852) [=Gammarus suluensis] B77
|--Ifalukia Barnard 1972 B77
| `--*I. resaca (Barnard 1965) [=Parelasmopus resacus] B77
|--Elasmopus Costa 1853 B77
| |--*E. rapax Costa 1853 B77
| |--E. affinis Della Valle 1893 BD95
| |--E. brasiliensis (Dana 1855) BD95
| |--E. pectenicrus (Bate 1862) BD95
| |--E. pocillimanus (Bate 1862) BD95
| `--E. rapax Costa 1853 BD95
|--Elasmopoides Stebbing 1908 B77
| `--*E. chevreuxi Stebbing 1908 B77
|--Beaudettia Barnard 1965 [Beaudettiidae] B77
| `--*B. palmeri Barnard 1965 B77
|--Cottesloe Barnard 1974 B77
| `--*C. berringar Barnard 1974 B77
|--Nuuana Barnard 1970 B77
| `--*N. amikai Barnard 1970 B77
|--Mallacoota Barnard 1972 B77
| `--*M. diemenensis (Haswell 1880) [=Megamoera diemenensis] B77
|--Maera B77
`--Quadrivisio Stebbing 1907 B77
|--*Q. bengalensis Stebbing 1907 B77
|--Q. chevreuxi Gordon 1968 S86
`--Q. lutzi (Shoemaker 1933) [=Pseudoceradocus lutzi; incl. Q. occidentalis Stephensen 1933] S86
*Type species of generic name indicated
REFERENCES
[BD95] Bakalem, A. & J.-C. Dauvin. 1995. Inventaire des crustacés amphipodes (Gammaridea, Caprellidea, Hyperiidea) des costes d'Algérie: essai de synthèse. Mésogée 54: 49-62.
[B77] Bousfield, E. L. 1977. A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Suppl. 4: 282-316.
[EHH05] Eberhard, S. M., S. A. Halse & W. F. Humphreys. 2005. Stygofauna in the Pilbara region, north-west Western Australia: a review. Journal of the Royal Society of Western Australia 88 (4): 167-176.
[FJK08] Finston, T. L., M. S. Johnson & B. Knott. 2008. A new genus and species of stygobitic paramelitid amphipod from the Pilbara, Western Australia. Records of the Western Australian Museum 24 (4): 395-410.
[K-SV02] Krapp-Schickel, T., & W. Vader. 2002. Redescription and generic position of Cheirocratella thori Stephensen, 1940 (Amphipoda, Gammaroidea). Sarsia 87: 9-18.
[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1-124.
[R86] Ruffo, S. 1986. Amphipoda: Groupe Metacrangonyx. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 550-552. E. J. Brill / Dr. W. Backhuys: Leiden.
[S86] Stock, J. H. 1986. Amphipoda: Melitid grouping (Melitidae sensu Bousfield, 1973, emend.) In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 504-513. E. J. Brill / Dr. W. Backhuys: Leiden.
[W86] Williams, W. D. 1986. Amphipoda on land-masses derived from Gondwana. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 553-559. E. J. Brill / Dr. W. Backhuys: Leiden.
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