Neocribellatae

Lampshade spider Hypochilus thorelli, copyright Marshal Hedin.

Belongs within: Araneida.
Contains: Argyronetidae, Hahniidae, Leptonetoidea, Plectana, Palpimanoidea, Eresoidea, Nicodamidae, Araneoidea, Austrochiloidea, Deinopoidea, Dictynidae, Amaurobioidea, Phyxelididae.

The Neocribellatae are a clade of spiders in which a sieve-like spinning plate, the cribellum, evolved in front of the spinnerets, though this feature has been secondarily lost in many derived subgroups. Members of this clade have a relatively short life cycle and do not moult subsequent to attaining maturity (Wunderlich 2019). Many members of this clade (most members of the subclade Praeentelegynae) have an entelegyne female reproductive system, with an epigyne bearing separate pairs of ducts for sperm transport during insemination and fertilisation, though it is debated whether such a system has arisen more than once.

<==Neocribellatae [Cleistospermiata] W19
    |  i. s.: Spinipalpus [Pholcochyroceridae] W19
    |           `--S. veta Wunderlich 2015 W19
    |--Hypochilidae [Palaearchaeidea, Paleocribellatae] W19
    |    |--Ectatosticta Simon 1892 [Ectatostictidae] JD-S07
    |    |    `--E. davidi A99
    |    `--Hypochilus Marx 1888 JD-S07
    |         |--H. coylei HR01
    |         |--H. gertschi FF99
    |         |--H. pococki FF99
    |         |--H. sheari HR01
    |         `--H. thorelli Marx 1888 HF08
    `--Praeentelegynae (see below for synonymy) W19
         |  i. s.: Argyronetidae S02
         |         Hahniidae S02
         |         Eurocoelotes inermis (Koch 1855) SL14
         |         Homalonychus Marx 1891 [Homalonychidae] JD-S07
         |           |--H. selenopoides R14
         |           `--H. theologus JD-S07
         |         Leptonetoidea W19
         |         Plectana D59
         |         Notocentria Thorell 1894 T94
         |           `--*N. sexspinosa Thorell 1894 T94
         |         Talthybia depressa Thorell 1898 GA11
         |--Palpimanoidea R14
         `--+--Eresoidea R14
            `--+--+--Nicodamidae R14
               |  `--+--Araneoidea R14
               |     `--+--Austrochiloidea R14
               |        `--+--Deinopoidea R14
               |           `--Dictynidae R14
               `--+--Amaurobioidea R14
                  `--Titanoecoidea JD-S07
                       |--Phyxelididae JD-S07
                       `--Titanoecidae JD-S07
                            |--Goeldia Keyserling 1891 JD-S07
                            |--Anuvinda Lehtinen 1967 JD-S07
                            |--Nurscia Simon 1874 JD-S07
                            |--Pandava Lehtinen 1967 JD-S07
                            `--Titanoeca Thorell 1869 R14, JD-S07
                                 |--T. americana R14
                                 `--T. obscura JD-S07

Praeentelegynae [Dictynoidea, Dipneumonomorpha, Entelegynae, Euetrioidae, Orbitelae, Orbiculariae, Orbitelariae] W19

*Type species of generic name indicated

REFERENCES

[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (transl. 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).

[D59] Doleschall, C. L. 1859. Tweede Bijdrage tot de kennis der Arachniden van den Indischen Archipel. Verhandelingen der Natuurkundige Vereeniging in Nederlandsch Indie [Acta Societatis Scientiarum Indo-Neêrlandicae] 5 (5): 1–60, pls 1–18.

[FF99] Forster, R., & L. Forster. 1999. Spiders of New Zealand and their World-wide Kin. University of Otago Press: Dunedin (New Zealand).

[GA11] Gregorič, M., I. Agnarsson, T. A. Blackledge & M. Kuntner. 2011. Darwin's bark spider: giant prey in giant orb webs (Caerostris darwini, Araneae: Araneidae)? Journal of Arachnology 39 (2): 287–295.

[HR01] Hale, R. J., & D. C. F. Rentz. 2001. The Gryllacrididae: an overview of the world fauna with emphasis on Australian examples. In: Field, L. H. (ed.) The Biology of Wetas, King Crickets and Their Allies pp. 95–110. CABI Publishing: Wallingford (UK).

[HF08] Huber, B. A., & N. Fleckenstein. 2008. Comb-hairs on the fourth tarsi in pholcid spiders. Journal of Arachnology 36 (2): 232–240.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[R14] Ramírez, M. J. 2014. The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History 390: 1–374.

[SL14] Samu, F., G. Lengyel, É. Szita, A. Bidló & P. Ódor. 2014. The effect of forest stand characteristics on spider diversity and species composition in deciduous-coniferous mixed forests. Journal of Arachnology 42 (2): 135–141.

[S02] Selden, P. A. 2002. Missing links between Argyroneta and Cybaeidae revealed by fossil spiders. Journal of Arachnology 30 (2): 189–200.

[T94] Thorell, T. 1894. Förteckning öfver Arachnider från Java och närgränsande öar, insamlade af docenten D:r Carl Aurivillius; jemte beskrifingar å några sydasiatiska och sydamerikanska Spindlar. Bihang till K. Svenska Vetenskaps-Akademiens Handlingar 20 pt. 4 (4): 1–63.

[W19] Wunderlich, J. 2019. What is a spider? Cretaceous fossils modify strongly phylogenetics as well as diagnoses of families, superfamilies and even suborders of spiders (Araneida) and other arthropods. Beiträge zur Araneologie 12: 1–32.