Basalhaplogynae

Desert bush spider Diguetia canities, from the National Park Service.

Belongs within: Araneida.
Contains: Scytodoidea, Pholcidae, Segestriidae, Dysderidae, Orsolobidae, Oonopidae.

The Basalhaplogynae are a clade of spiders in which the chelicerae are fused at the base and provided with a lamina; members of the clade also have the tegulum and subtegulum fused and the spinnerets lack tartipores (Jocqué & Dippenaar-Schoeman 2007).

<==Basalhaplogynae [Haplogynae, Pholcoidea] W19
    |--Filistatidae [Filistatomorpha] JD-S07
    |    |  i. s.: Misionella P05
    |    |           |--M. didicostae Penney 2005 P05
    |    |           `--M. mendensis R14
    |    |         Microfilistata Zohnstein 1990 G95
    |    |--Filistatinae G95
    |    |    |  i. s.: Zaituna Lehtinen 1967 G95
    |    |    |--Sahastata Benoit 1968 G95
    |    |    |    |--S. nigra G95
    |    |    |    `--S. sabaea Brignoli 1982
    |    |    `--+--Kukulcania Lehtinen 1967 G95
    |    |       |    `--K. hibernalis G95
    |    |       `--Filistata Latreille 1810 G95, JD-S07
    |    |            |--F. hibernalis B06
    |    |            `--F. insidiatrix (Forskål 1775) MG03
    |    `--Prithinae G95
    |         |  i. s.: Malalistata Mello-Leitão 1946 G95
    |         |         Tricalamus Wang 1987 G95
    |         |--+--Filistatinella Gertsch & Ivie 1936 G95
    |         |  `--+--Filistatoides P.-Cambridge 1899 G95
    |         |     `--Pikelinia Mello-Leitão 1946 G95
    |         |          `--P. tambilloi R14
    |         `--+--+--Andoharano Lehtinen 1967 G95
    |            |  `--Afrofilistata Benoit 1968 G95
    |            `--+--Pritha Lehtinen 1967 G95, MG03
    |               |    |--*P. nana (Simon 1868) MG03
    |               |    `--P. crosbyi (Spassky 1938) [=Filistata crosbyi] MG03
    |               `--+--Yardiella Gray 1994 G95
    |                  `--Wandella Gray 1994 G95
    |                       |--W. orana G95
    |                       `--W. waldockae MC13
    `--Synspermiata [Scytodiformes] W19
         |--+--Scytodoidea JD-S07
         |  `--+--Pholcidae JD-S07
         |     `--+--Plectreuridae JD-S07
         |        |    |--Kibramoa Chamberlin 1924 JD-S07
         |        |    `--Plectreurys Simon 1893 JD-S07
         |        |         `--P. tristis Simon 1893 HF08
         |        `--Diguetidae JD-S07
         |             |--Segestrioides Keyserling 1883 JD-S07
         |             |    `--S. bicolor JD-S07
         |             `--Diguetia Simon 1895 JD-S07
         |                  |--D. canities JD-S07
         |                  `--D. signata Gertsch 1958 HF08
         `--+--+--Dysderoidea JD-S07
            |  |    |  i. s.: Trogloraptor [Trogloraptoridae] R14
            |  |    |--Segestriidae JD-S07
            |  |    `--+--Dysderidae JD-S07
            |  |       `--+--Orsolobidae JD-S07
            |  |          `--Oonopidae JD-S07
            |  `--Tetrablemmidae JD-S07
            |       |--Shearella JD-S07
            |       |--Afroblemma JD-S07
            |       |--Tetrablemma P.-Cambridge 1873 [Tetrablemminae] JD-S07
            |       |--Paculla [Pacullinae] JD-S07
            |       `--Monoblemma spinosum Wunderlich 1988 S93
            `--Caponiidae JD-S07
                 |--Caponia Simon 1887 [Caponiinae] JD-S07
                 |--Calponia Platnick 1993 JD-S07
                 |--Diploglena Purcell 1904 JD-S07
                 |--Nopsides Chamberlin 1924 JD-S07
                 |--Notnops Platnick 1994 JD-S07
                 |--Orthonops Chamberlin 1924 JD-S07
                 |--Taintnops Platnick 1994 JD-S07
                 |--Tarsonops Platnick 1994 JD-S07
                 |--Tisentnops Platnick 1994 JD-S07
                 |--Caponina Simon 1887 JD-S07
                 |    `--C. notabilis (Mello-Leitão 1939) FM11
                 `--Nops MacLeay 1839 [Nopinae] JD-S07
                      |--N. lobatus Wunderlich 1988 S93
                      `--N. segmentatus Wunderlich 1988 S93

*Type species of generic name indicated

REFERENCES

[B06] Banks, N. 1906. Arachnida from the Bahamas. Bulletin of the American Museum of Natural History 22: 185–189.

[FM11] Freire-Jr, G. de B., & P. C. Motta. 2011. Effects of experimental fire regimes on the abundance and diversity of cursorial arachnids of Brazilian savannah (cerrado biome). Journal of Arachnology 39 (2): 263–272.

[G95] Gray, M. R. 1995. Morphology and relationships within the spider family Filistatidae (Araneae: Araneomorphae). Records of the Western Australian Museum Supplement 52: 79–89.

[HF08] Huber, B. A., & N. Fleckenstein. 2008. Comb-hairs on the fourth tarsi in pholcid spiders. Journal of Arachnology 36 (2): 232–240.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[MG03] Marusik, Yu. M., & E. F. Guseinov. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 1. New family and genus records. Arthropoda Selecta 12 (1): 29–46.

[P05] Penney, D. 2005. Fossil blood droplets in Miocene Dominican amber yield clues to speed and direction of resin secretion. Palaeontology 48 (5): 925–927.

[R14] Ramírez, M. J. 2014. The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History 390: 1–374.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[W19] Wunderlich, J. 2019. What is a spider? Cretaceous fossils modify strongly phylogenetics as well as diagnoses of families, superfamilies and even suborders of spiders (Araneida) and other arthropods. Beiträge zur Araneologie 12: 1–32.