Holcobunus nigripalpis, from here.

Belongs within: Sclerosomatidae.

Holcobunus is a genus of sclerosomatid harvestmen found in southern Brazil, characterised by a sinuous penis bearing an elongate stylus.

Characters (from Tourinho et al. 2015): Pedipalpal femur, patella and proximal half of tibia black. Eyemound dorsally armed with granules or spines with points either sharp or blunt, located on its anterior half or arranged in two parallel rows. Lateral margins of supracheliceral lamina crenulate. Dorsal scute in male trapezoidal with posterior margin convex, in female markedly pyriform. Pedipalpal tibia sinuous at base. Femoral pseudoarticular formula: 2/4–6/2–3/2–4. Winglets of penis uniformly narrow (longer than wide) along its length, borders undulated; anterior part in ventral view folding ventrally to form collar. Shaft strongly sinuous dorsoventrally, slightly bent without forming definite angle at glans. Glans globose, surface with simple pores and two or three pairs of asymmetric apical setae, stylus 40–70% of glans length.

<==Holcobunus Roewer 1910 [incl. Paratamboicus Mello-Leitão 1940, Sympathica Mello-Leitão 1933] TK01
    |--*H. nigripalpis Roewer 1910 (see below for synonymy) TK01
    |--‘Sympathica’ aureopunctata Roewer 1953 TK01
    |--H. bicornutus (Mello-Leitão 1940) [=*Paratamboicus bicornutus] TK01
    |--H. dissimilis Tourinho, Pinto-da-Rocha & Bragagnolo 2015 TP-RB15
    |--H. dubius Ringuelet 1959 T-D04
    |--‘Sympathica’ formosa Ringuelet 1953 TK01
    |--H. ibitirama Tourinho, Pinto-da-Rocha & Bragagnolo 2015 TP-RB15
    |--H. iguassuensis Mello-Leitão 1935 TK01
    |--H. luteipalpis Roewer 1910 TK01
    |--H. marmoratus Mello-Leitão 1938 (n. d.) TK01
    |--H. missionicus Ringuelet 1959 T-D04
    |--H. segadasi Mello-Leitão 1949 (n. d.) TK01
    |--H. tocantinus Roewer 1953 KP-R02
    |--H. trochanteralis Roewer 1953 KP-R02
    |--H. uaisoh Tourinho, Pinto-da-Rocha & Bragagnolo 2015 TP-RB15
    |--H. unicolor (Loman 1902) [=Prionomma unicolor] R23
    `--H. unifasciatus Roewer 1910 R23

*Holcobunus nigripalpis Roewer 1910 [incl. *Sympathica bisignata Mello-Leitão 1933, S. inermis Mello-Leitão 1933 (n. n.), Tamboicus silvicola Mello-Leitão 1932, S. silvicola] TK01

*Type species of generic name indicated


[KP-R02] Kury, A. B., & R. Pinto-da-Rocha. 2002. Opiliones. In: Adis, J. (ed.) Amazonian Arachnida and Myriapoda pp. 345–362. Pensoft: Sofia.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[TK01] Tourinho, A. L. M., & A. B. Kury. 2001. Notes on Holcobunus Roewer, 1910, with two new generic synonymies (Arachnida, Opiliones, Sclerosomatidae). Boletim do Museu Nacional, nova serie, Zoologia 461: 1–22.

[TP-RB15] Tourinho, A. L., R. Pinto-da-Rocha & C. Bragagnolo. 2015. Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425–436.

[T-D04] Tourinho-Davis, A. L. 2004. A new genus of Gagrellinae from Brazil, with a comparative study of some of the subtropical and southernmost tropical South American genera (Opiliones, Eupnoi, Sclerosomatidae). Revista Ibérica de Aracnología 9: 157–177.

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