Dorsal views of female (left) and male of Harmanda instructa aenescens, from Martens (1987).

Belongs within: Sclerosomatidae.

Harmanda is a genus of sclerosomatid harvestmen primarily known from the Himalayan region; species assigned to this genus from other parts of Asia may be misplaced (Martens 1987).

See also: More Gagrellinae (Taxon of the Week: Harmanda).

Characters (from Martens 1987): Shaft of penis strongly dorsoventrally flattened, narrow, largely paralell-sided, without a demarcated basis, muscles occupying at least half of shaft. Dorsal mantle of pockets well developed, mostly significantly overlapping ventrally, mantle often with pigment; distinct wings along margin of shaft, often nearly reaching nearly ventral opening, rarely completely missing. Pedipalps slender, apophyses strongly developed, especially on patella, less so on tibia, weaker in female than male. Sexual dimorphism clearly present in some species in dorsal colour patterning or gland formation on tibia of male pedipalp.

<==Harmanda Roewer 1910 [=Harmandella Roewer 1910; incl. Ischnobunus Roewer 1955] C92
    |--*H. instructa Roewer 1910 [=*Harmandella instructa] C92
    |    |--H. i. instructa M87 [incl. Harmandella lineata Roewer 1911 M87, Harmanda lineata R55]
    |    |--H. i. aenescens (Roewer 1911) [=Harmandella aenescens; incl. Adungrella atrorubra Suzuki 1967] M87
    |    |--H. i. bhutanensis Martens 1987 M87
    |    `--H. i. lineatocoxa Martens 1987 M87
    |--H. annulata (Roewer 1911) R55 [=Harmandella annulata R23]
    |--H. arunensis Martens 1987 M87
    |--H. beroni Martens 1987 M87
    |--H. caerulea (Roewer 1910) [=Zaleptus caeruleus; incl. Z. splendens Roewer 1911] M87
    |--H. corrugata Martens 1987 M87
    |--H. elegantula (Roewer 1955) [=*Ischnobunus elegantulus] C92
    |--H. kanoi (Suzuki 1967) [=Verpulus kanoi] M87
    |--H. khumbua Martens 1987 M87
    |--H. latephippiata Martens 1987 M87
    |--H. medioimmicans Martens 1987 M87
    |--H. nigrolineata Martens 1987 M87
    `--H. triseriata Roewer 1923 R55

*Type species of generic name indicated


[C92] Crawford, R. L. 1992. Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History 8: 1–60.

[M87] Martens, J. 1987. Opiliones aus dem Nepal-Himalaya. VI. Gagrellinae (Arachnida: Phalangiidae). Courier Forschungsinstitut Senckenberg 93: 87–202.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[R55] Roewer, C. F. 1955. Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 4. Teil (Schluß). Senckenbergiana Biologica 36 (3–4): 123–171.

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