Gagrella amboinensis, copyright Michael Kesl.

Belongs within: Sclerosomatidae.

Gagrella is a diverse genus of sclerosomatid harvestmen recognised from southern Asia and northern Australasia, currently lacking a properly defined diagnosis and hence of uncertain monophyly.

Characters (from Martens 1987): Femur II only with a single pseudoarticular nodule; area 2 or areas 1 and 2 of scutum each with a median cone; eyemound low, smooth or granular, without large spines; femora I and III cylindrical, longer than body.

<==Gagrella Stoliczka 1869 T09 [incl. Crassicippus Roewer 1910 C92]
    |--*G. signata Stoliczka 1869 T09 [=Crassicippus signatus B30]
    |--G. aenescens Thorell 1889 S85
    |--G. aenigra Roewer 1954 R54
    |--G. alba Suzuki 1985 S85
    |--G. albertisii Thorell 1876 T09 (see below for synonymy)
    |--G. albicoxata Roewer 1954 R54
    |--G. albifrons Roewer 1915 R54
    |--G. amboinensis (Doleschall 1857) T09 [=Phalangium amboinense C92; incl. Gagrella mediocris Kulczynski 1910 R23]
    |    |--G. a. amboinensis T09
    |    `--G. a. bicornuta Strand 1910 T09
    |--G. andamana Roewer 1929 R54
    |--G. annapurnica Martens 1987 M87
    |--G. annulatipes Roewer 1912 R54
    |--G. aorana Roewer 1954 R54
    |--G. argentea Roewer 1913 R54
    |--G. armillata Thorell 1889 R54
    |--G. atrorubra Simon 1901 R54
    |--G. aura (Rainbow 1913) T09 [=Liobunum aurum R23]
    |--G. aureolata Roewer 1915 R54
    |--G. aurispina Roewer 1954 R54
    |--G. auromaculata Roewer 1954 R54
    |--G. auroscutata Roewer 1954 R54
    |--G. bakeri Roewer 1926 R54
    |--G. beauforti Roewer 1913 R54
    |--G. bella Roewer 1954 R54
    |--G. bengalica Roewer 1954 R54
    |--G. biarmata Roewer 1954 R54
    |--G. bicolorispina Roewer 1954 R54
    |--G. binotata Simon 1887 R54
    |--G. biseriata Simon 1901 R54 [incl. G. illusa Simon 1902 R23]
    |--G. borneoensis Roewer 1954 R54
    |--G. brunnea Roewer 1954 R54
    |--G. buruana Roewer 1954 R54
    |--G. caerulea Roewer 1910 R54
    |--G. cana Roewer 1954 S85
    |--G. carinia Roewer 1936 R54
    |--G. cauricrepa Taylor 2009 T09
    |--G. ceramensis Roewer 1913 R54
    |    |--G. c. ceramensis R54
    |    `--G. c. ‘bispinosa’ Roewer 1913 non G. feae bispinosa Thorell 1889 R54
    |--G. cerata Roewer 1913 R54
    |--G. cervina Simon 1887 R54
    |--G. cinerascens Roewer 1910 R54
    |--G. concinna Thorell 1891 R54
    |--G. coriacea Roewer 1910 S85
    |--G. corrugata Roewer 1954 R54
    |--G. crassitarsis Suzuki 1969 S85
    |--G. cruciata Roewer 1923 R54
    |--G. cuprea Roewer 1910 R54
    |--G. cuprinitens Roewer 1954 R54
    |--G. curvispina Roewer 1913 R54
    |--G. cyanargentea Roewer 1915 S85
    |--G. cyanatra Roewer 1954 R54
    |--G. delicatula Roewer 1954 R54
    |--G. denticulatifrons Roewer 1954 T09
    |--G. disticta Thorell 1889 S85 [=G. nocticolor var. disticta R23]
    |--G. docilis W03
    |--G. dubia Giltay 1930 T09
    |--G. duplex Roewer 1915 T09
    |--G. elegans Simon 1877 R54
    |--G. ephippiata Thorell 1891 [=Crassicippus ephippiatus] R23
    |--G. erebea Thorell 1889 R54
    |--‘Crassicippus’ figuratus Roewer 1923 R23
    |--G. flava Roewer 1910 R54
    |--G. flavimaculata With 1903 R54
    |--G. fokiensis Roewer 1954 R54
    |--G. foveolata Roewer 1923 R54
    |--G. franzi Martens 1987 M87
    |--G. fulva Roewer 1910 S85
    |--G. fumosa Suzuki 1985 S85
    |--G. fuscipes Roewer 1910 T09
    |--G. gilva Roewer 1926 R54
    |--G. grandis Roewer 1910 R54
    |--G. granobunus Roewer 1954 T09
    |--G. gravelyi Roewer 1912 R54
    |--G. grisea Roewer 1911 R54
    |--G. hainanensis Roewer 1911 R54
    |--G. hasseltii Thorell 1891 R54
    |--G. heinrichi Roewer 1954 R54
    |--G. histrionica Thorell 1889 R54
    |--G. indochinensis Roewer 1927 R54
    |--G. infuscata Roewer 1911 R54
    |--G. insculpta Pocock 1897 R54 [=Hexomma insculpta B30]
    |--G. insulana Roewer 1954 R54
    |--G. iwamasai Suzuki 1977 S77
    |--G. lateritia Roewer 1954 R54
    |--G. leopoldi Giltay 1930 T09
    |--G. lepida Thorell 1889 S85
    |    |--G. l. lepida R54
    |    |--G. l. claducha Thorell 1889 R54
    |    `--G. l. dibelona Thorell 1889 R54
    |--G. leucobunus Roewer 1912 R54
    |--G. lineatipes Roewer 1911 R54
    |--G. longipalpis Thorell 1891 R54 [=Hexomma longipalpis B30]
    |--G. longipes Suzuki 1982 S85
    |--G. longispina Roewer 1913 R54
    |--G. luteofrontalis Roewer 1910 S85
    |--G. luzonica Loman 1902 R54
    |--G. magnifica Roewer 1910 R54
    |--G. malabarica Roewer 1954 R54
    |--G. marginata Roewer 1954 R54
    |--G. mertoni Strand 1911 R54
    |--G. metallica Roewer 1929 R54
    |--G. minuta Roewer 1954 R54
    |--G. mjobergi (Banks 1930) [=Hexomma mjobergi] R54
    |--G. modesta Suzuki 1985 S85
    |--G. monacantha (Herbst 1798) (n. d.) [=Opilio monacantha, Phalangium monacanthum] R23
    |--G. monticola Thorell 1891 R54
    |    |--G. m. monticola R54
    |    `--G. m. tarda Thorell 1891 R54
    |--G. natuna Roewer 1912 R54
    |--G. neocera Forster 1949 T09
    |--‘Crassicippus’ nigerrimus Roewer 1910 R23
    |--G. nigrescens Roewer 1954 R54
    |--G. nigripalpis Roewer 1910 R54
    |--G. niponica Roewer 1954 R54
    |--G. nobilis With 1903 R54
    |--G. nocticolor Thorell 1889 (n. d.) R23
    |--G. obscura Simon 1877 R54
    |--G. ochroleuca Roewer 1954 R54
    |--G. opaca Roewer 1954 R54
    |--G. ornata Roewer 1910 R54
    |--G. palnica Roewer 1929 R54
    |--G. parallela Roewer 1931 R54
    |--G. parva Roewer 1929 R54
    |--G. patalungensis Simon 1901 R54
    |--G. paupera With 1905 R54 [=Eugagrella paupera R23]
    |--G. prasina Roewer 1911 R54
    |--G. pretiosa (Banks 1930) [=Hexomma pretiosa] R54
    |--G. promeana Roewer 1954 T09
    |--G. pulcra Suzuki 1985 S85
    |--G. pullata Thorell 1891 S85
    |--G. punctata Roewer 1954 R54
    |--G. quadrimaculata Roewer 1954 S85
    |--G. quadriseriata Roewer 1954 R54
    |--G. quadrivittata Simon 1887 [=Crassicippus quadrivittatus] R23
    |--G. rorida Roewer 1954 R54
    |--G. rufa Roewer 1954 R54
    |--G. sadona Roewer 1954 R54
    |--G. sampitia Roewer 1954 R54
    |--G. sarasinorum Roewer 1913 R54
    |--G. sarawakensis With 1905 R54 [=Hexomma sarawakensis B30]
    |--G. satarana Roewer 1954 R54
    |--G. scabra Roewer 1912 R54
    |--G. scintillans Roewer 1910 R54
    |--G. scrobiculata Thorell 1891 R54 [=Hexomma scrobiculata B30]
    |--G. scutaurea Roewer 1954 R54
    |--G. semigranosa Simon 1901 [=*Crassicippus semigranosus] C92
    |--G. serrulata Roewer 1910 R54
    |--G. sexmaculata Suzuki 1969 S85
    |--G. sherriffsi Roewer 1954 R54
    |--G. sinensis Roewer 1954 R54
    |--G. speciosa Roewer 1911 R54
    |--‘Crassicippus speciosus’ Roewer 1912 non Gagrella speciosa Roewer 1911 R23
    |--G. spinoculata Roewer 1931 R54
    |--G. spinulosa Thorell 1889 S85
    |--G. subfusca Roewer 1910 R54
    |--G. sulphurea Roewer 1912 S85
    |--G. suluana Roewer 1954 R54
    |--G. sumba Roewer 1954 R54
    |--G. testacea Roewer 1954 non Phalangium testaceum Doleschall 1859 (not preoc. if dif. gen.) R54
    |--G. thaiensis Suzuki 1982 S85
    |--G. thorelli (Banks 1930) [=Hexomma thorelli] R54
    |--G. tibialis Roewer 1931 S85
    |--G. tinjurae Martens 1987 M87
    |--G. tricolor Roewer 1954 R54
    |--G. tuberculata Roewer 1913 R54
    |--G. turki Roewer 1954 R54
    |--G. unispinosa With 1903 R54 [=G. imperator unispinosa R23]
    |--G. victoria Roewer 1954 R54
    |--G. viridalba Roewer 1954 R54
    |--G. viridargentea Roewer 1954 R54
    |--G. viridula Roewer 1929 R54
    `--G. yodoensis Roewer 1954 R54

Gagrella albertisii Thorell 1876 T09 [incl. G. aruella Strand 1911 R23, G. bipeltata Thorell 1891 R23, G. doleschallii Thorell 1876 R23, G. bipeltata var. inquirenda Strand 1910 S10, R23, G. samangana Strand 1911 R23, G. xanthostoma Thorell 1891 R23]

*Type species of generic name indicated


[B30] Banks, N. 1930. Phalangida from Borneo. Sarawak Museum Journal, new series 4 (1): 57–86, pls 11–12.

[C92] Crawford, R. L. 1992. Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History 8: 1–60.

[M87] Martens, J. 1987. Opiliones aus dem Nepal-Himalaya. VI. Gagrellinae (Arachnida: Phalangiidae). Courier Forschungsinstitut Senckenberg 93: 87–202.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[R54] Roewer, C. F. 1954. Indoaustralische Gagrellinae (Opiliones, Arachnidae). (Weitere Weberknechte XVIII). 1. Teil. Senckenbergiana Biologica 35 (3–4): 181–236.

[S10] Strand, E. 1910. Spinnentiere aus Neuguinea (Opiliones, Psechridae und Clubionidae) gesammelt von Dr. Schlaginhaufen. Abhandlungen und Berichte des K├Ânigl. Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 13 (5): 1–16.

[S77] Suzuki, S. 1977. Opiliones from Taiwan (Arachnida). Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology) 27 (1): 121–157.

[S85] Suzuki, S. 1985. A synopsis of the Opiliones of Thailand (Arachnida) II. Palpatores. Steenstrupia 11 (7): 209–257.

[T09] Taylor, C. K. 2009. Revision of the Australian Gagrellinae (Arachnida: Opiliones: Sclerosomatidae), with a description of a new species. Australian Journal of Entomology 48: 217–222.

[W03] With, C. 1903. New and old Phalangiidae from the Indian region. Journal of the Linnean Society, Zoology 28: 466–509.

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