Phyllodocidae

Hesionura elongata, copyright Hans Hillewaert.

Belongs within: Phyllodocida.

The Phyllodocidae are a group of active marine worms with leaf-shaped cirri.

Characters (from Howell 1962): Body long and narrow, with many segments bearing one, two, or more setae; dorsal and ventral cirri leaf-shaped; two anal cirri may be present.

<==Phyllodocidae
    |  i. s.: Palaeochaeta Clarke 1903 H62
    |           `--*P. devonica Clarke 1903 H62
    |         Levisettius campylonectus PTV14
    |--+--+--Sige NP10
    |  |  |    |--S. fusigera NP10
    |  |  |    `--S. limbata M62
    |  |  `--Eulalia viridis (Linnaeus 1767) NP10, RP07
    |  `--+--Pseudomystides limbata NP10
    |     |--+--‘Eulalia’ bilineata NP10
    |     |  `--Protomystides exigua NP10
    |     `--+--‘Eulalia’ mustela NP10
    |        `--+--Pterocirrus macroceros NP10 [=Eulalia macroceros K92]
    |           `--+--Eumida NP10
    |              |    |--E. arctica NP10
    |              |    |--E. sanquinea KBC03
    |              |    `--E. tubiformis KBC03
    |              `--Alciopidae NP10
    |                   |--Torrea NP10
    |                   `--Alciopina NP10
    `--+--+--Chaetoparia nilssoni NP10
       |  `--Notophyllum foliosum (Sars 1835) NP10, RP07
       `--+--Phyllodoce Ranzani 1817 NP10, H62
          |    |--*P. maxillosa Ranzani 1817 H62
          |    |--P. groenlandica (Oersted 1843) SS07
          |    |--P. longipes NP10
          |    |--P. mucosa Oersted 1842 A70
          |    `--P. vittata Ehlers 1864 A70
          `--+--Nereiphylla lutea NP10
             `--Paranaitis NP10
                  |--P. wahlbergi NP10
                  `--+--P. kosteriensis NP10
                     `--Eteoninae H-S86
                          |--Eteone NP10
                          |    |--E. longa (Fabricius 1780) RP07
                          |    `--E. picta Quatrefages 1865 RP07
                          `--Hesionura Hartmann-Schröder 1958 H-S86
                               |--H. augeneri (Friedrich 1937) H-S86
                               |--H. coineaui (Laubier 1962) H-S86
                               |    |--H. c. coineaui H-S86
                               |    `--H. c. difficilis (Banse 1963) H-S86
                               |--H. elongata (Southern 1914) H-S86
                               |--H. fragilis Hartmann-Schröder 1958 H-S86
                               |--H. japoni Yamanishi 1980 H-S86
                               |--H. laubieri Hartmann-Schröder 1963 H-S86
                               |--H. mystidoides (Hartmann-Schröder 1963) H-S86
                               |--H. natalensis Hartmann-Schröder 1974 H-S86
                               |--H. portmanni Laubier 1967 H-S86
                               `--H. serrata (Hartmann-Schröder 1960) H-S86

*Type species of generic name indicated

REFERENCES

[A70] Amoureux, L. 1970. Annélides Polychètes du Bassin de chasse d’Ostende: inventaire d’une petite collection. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 46 (6): 23–24.

[H-S86] Hartmann-Schröder, G. 1986. Polychaeta (incl. Archiannelida). In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 210–233. E. J. Brill/Dr W. Backhuys: Leiden.

[H62] Howell, B. F. 1962. Worms. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt W. Miscellanea: Conodonts, Conoidal Shells of Uncertain Affinities, Worms, Trace Fossils and Problematica pp. W144–W177. Geological Society of America, and University of Kansas Press.

[KBC03] Kashin, I. A., E. V. Bagaveeva & S. F. Chaplygina. 2003. Fouling communities of hydrotechnical constructions in Nakhodka Bay (Sea of Japan). Russian Journal of Marine Biology 29: 267–283.

[K92] Kozloff, E. N. 1992. The genera of the phylum Orthonectida. Cahiers de Biologie Marine 33: 377–406.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[NP10] Nygren, A. & F. Pleijel. 2010. Chimaeras and the origins of the holopelagic annelids Typhloscolecidae and Lopadorhynchidae: a reply to Struck and Halanych (2010). Zoologica Scripta 40 (1): 112–114.

[PTV14] Parry, L., A. Tanner & J. Vinther. 2014. The origin of annelids. Palaeontology 57 (6): 1091–1103.

[RP07] Rousset, V., F. Pleijel, G. W. Rouse, C. Erséus & M. E. Siddall. 2007. A molecular phylogeny of annelids. Cladistics 23: 41–63.

[SS07] Struck, T. H., N. Schutt, T. Kusen, E. Hickman, C. Bleidorn, D. McHugh & K. M. Halanych. 2007. Annelid phylogeny and the status of Sipuncula and Echiura. BMC Evolutionary Biology 7: 57.