Italian bugloss Anchusa azurea, copyright Alberto Salguero Quiles.

Belongs within: Boraginaceae.

The Boragineae is a group of plants in the family Boraginaceae found almost entirely in Eurasia, characterised by flowers with a simple style and usually well-developed faucal appendages (Långström & Chase 2002).

Characters (from Långström & Chase 2002): Flowers with usually well-developed faucal appendages; style simple, with one or two stigmas; pollen colporate; nutlets with basal scar often surrounded by annular rim, gynobase flat.

<==Boragineae [Anchusae, Borageae]
    |--Borago officinalis Linnaeus 1753 LC02, CD07
    `--+--+--Nonea versicolor LC02
       |  `--Pentaglottis sempervirens LC02
       `--Anchusa LC02
            |--A. azurea H91
            |--A. caespitosa SL06
            |--A. capensis BR65
            |--A. officinalis LC02
            |--A. ovata O88
            `--A. spathulata C45

Boragineae incertae sedis:
  Anchusella LC02
  Brunnera LC02
  Caryolopha sempervirens LC02, C55b [=Anchusa sempervirens C55b]
  Cynoglottis barrelieri LC02, GR98
  Elizaldia LC02
  Oskampia LC02
  Psilostemon LC02
  Stomotechium LC02
  Trachystemon LC02
  Symphytum LC02
    |--S. asperrimum C55b
    |--S. caucasicum V09
    |--S. cordatum H09
    `--S. officinale C55b
  Lycopsis LC02
    |--L. arvensis [=Anchusa arvensis] BR65
    |--L. orientalis C55b
    `--L. variegata C55a
  Pulmonaria LC02
    |--P. mollis C55a
    |--P. officinalis C55b
    |--P. rubra H09
    `--P. virginica C55b

*Type species of generic name indicated


[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae-Compositae. W. L. Hawes, Government Printer: Adelaide.

[C55a] Candolle, A. de. 1855a. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 1. Librairie de Victor Masson: Paris.

[C55b] Candolle, A. de. 1855b. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[CD07] Cantino, P. D., J. A. Doyle, S. W. Graham, W. S. Judd, R. G. Olmstead, D. E. Soltis, P. S. Soltis & M. J. Donoghue. 2007. Towards a phylogenetic nomenclature of Tracheophyta. Taxon 56 (3): E1-E44.

[C45] Colenso, W. 1845. Memoranda of an excursion, made in the Northern Island of New Zealand, in the summer of 1841-2; intended as a contribution towards the natural productions of the New Zealand groupe: with particular reference to their botany. Tasmanian Journal of Natural Science 2: 210-234.

[GR98] Greuter, W., & T. Raus (eds.) 1998. Med-Checklist Notulae, 17. Willdenowia 28: 163-174.

[H09] Heltmann, H. 2009. Der Königstein (Piatra Craiului), die Perle der Burzenländer Gebirge. Mauritiana 20 (3): 515-527.

[H91] Hubálek, Z. 1991. Biogeographic indication of natural foci of tick-borne infections. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 255-260. SPB Academic Publishing: The Hague.

[LC02] Långström, E., & M. W. Chase. 2002. Tribes of Boraginoideae (Boraginaceae) and placement of Antiphytum, Echiochilon, Ogastemma and Sericostoma: a phylogenetic analysis based on atpB plastid DNA sequence data. Plant Systematics and Evolution 234: 137-153.

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: An introductory note. In The Himalayan Plants vol. 1 (H. Ohba & S. B. Malla, eds) The University Museum, University of Tokyo, Bulletin 31: 19-46.

[SL06] Schulz, H.-J., & P. Lymberakis. 2006. First contribution to the knowledge of the Collembola fauna of the White Mountains (Lefká Óri) in west Crete (Insecta, Collembola, Isotomidae). Senckenbergiana Biologica 86 (2): 229-234.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183-194.

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