Arran brown Erebia ligea, photographed by Harald Süpfle.

Belongs within: Satyrinae.
Contains: Euptychia.

The Satyrini are a diverse group of butterflies with larvae that feed mostly on grasses.

    |  i. s.: Heteronympha WWN03
    |           |--H. cordace NS04
    |           |    |--H. c. cordace NS04
    |           |    `--H. c. wilsoni NS04
    |           `--H. merope WWN03
    |         Cercyonis pegala WWN03
    |         Aphantopus hyperanthus WWN03
    |--Hypocystina J03
    |    |--Zipaetis J03
    |    |--Lamprolenis J03
    |    |--Hypocysta J03
    |    `--Xenica J03
    |--+--Dirina J03
    |  |--Melanargiina J03
    |  `--+--Satyrina J03
    |     `--Pronophilina J03
    `--+--+--Maniola WWN03 [Maniolina J03]
       |  |    `--M. jurtina WWN03
       |  `--Coenonympha B66 [Coenonymphina J03]
       |       `--C. oedipus [=Neonympha oedipus] B66
       |            |--C. o. oedipus I92
       |            `--C. o. annulifer Butler 1877 I92
       |--Erebia B66 [Erebiina J03]
       |    |--‘Papilio’ florimel Fabricius 1793 [=Neonympha florimel, Satyrus florimel] B66
       |    |--E. ligea I92
       |    |    |--E. l. ligea I92
       |    |    |--E. l. rishirizana Matsumura 1928 I92
       |    |    `--E. l. takanonis Matsumura 1909 I92
       |    |--E. niphonica I92
       |    `--E. palarica WWN03
       `--+--Euptychia J03
          `--Ypthimina J03
               |--Austroypthima J03
               `--Ypthima [incl. Xois] J03
                    |--‘Xois’ fulvida PP02
                    |--Y. masakii Ito 1947 I92
                    |--Y. riukiuana Matsumura 1906 I92
                    |--‘Xois’ sesara PP02
                    `--Y. yayeyamana Nire 1920 I92

*Type species of generic name indicated


[B66] Butler, A. G. 1866. A monograph of the genus Euptychia, a numerous race of butterflies belonging to the family Satyridae; with descriptions of sixty species new to science, and notes on their affinities &c. Proceedings of the Zoological Society of London 1866: 458-504.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[J03] Jong, R. de. 2003. Are there butterflies with Gondwanan ancestry in the Australian region? Invertebrate Systematics 17: 143-156.

[NS04] New, T. R., & D. P. A. Sands. 2004. Management of threatened insect species in Australia, with particular reference to butterflies. Australian Journal of Entomology 43 (3): 258-270.

[PP02] Patrick, B., & H. Patrick. 2002. Butterflies of Fiji. Weta 24: 5-12.

[WWN03] Wahlberg, N., E. Weingartner & S. Nylin. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 28: 473-484.

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