Northern crescent Phyciodes cocyta, photographed by Gilles Gonthier.

Belongs within: Nymphalinae.

The Melitaeini include the checkerspots, a primarily Nearctic group of butterflies. Checkerspots generally have wings marked with red, orange and black or brown. The genus Melitaea, fritillaries, contains about 80 species found in the Palaearctic region.

    |--Euphydryas WWN03
    |    |--E. aurinia [=Melitaea aurinia] G91
    |    |--E. chalcedona D37
    |    |--E. desfontainii WWN03
    |    `--E. editha S85
    |         |--E. e. editha D37
    |         `--E. e. wrightii D37
    `--+--Chlosyne WWN03
       |    |--C. janais Drury 1782 FS54
       |    `--C. lacinia (Geyer 1837) FS54
       `--+--Melitaea WWN03
          |    |--M. arcesia P01
          |    |--M. athalia [incl. M. athalia ab. navarina] P01
          |    |--M. cinxia P01
          |    |--M. didyma [incl. M. didyma var. alpina] P01
          |    |--M. didymoides WWN03
          |    |--M. neumoegeni D37
          |    |--M. scotosia Butler 1878 I92
          |    |--M. theona (Ménétriés 1857) FS54
          |    `--M. wrightii D37
          `--Phyciodes WWN03
               |--P. albescens Röber 1913 FS54
               |--P. cocyta WWN03
               |--P. elaphiaea Hewitson 1868 S55
               |--P. liriope S55
               |    |--P. l. liriope S55
               |    `--P. l. fragilis (Bates 1864) S55
               `--P. perilla S55
                    |--P. p. perilla S55
                    `--P. p. lugubris Röber 1913 S55

*Type species of generic name indicated


[D37] Dobzhansky, T. 1937. Genetics and the Origin of Species. Columbia University Press: New York.

[FS54] Franz, E., & H. Schröder. 1954. Tagfalter (Lep. Rhopalocera) aus El Salvador. Senckenbergiana Biologica 35: 75–87.

[G91] Groves, C. P. 1991. A Theory of Human and Primate Evolution, revised ed. Clarendon Press: Oxford.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[P01] Pável, J. 1901. Lepkék [Lepidopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 171–177. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[S55] Schröder, H. 1955. Eine Falter-Ausbeute aus dem westlichen Bolivien. (Ins. Lepid. Rhopal.) Senckenbergiana Biologica 36: 329–338.

[S85] Slatkin, M. 1985. Gene flow in natural populations. Annual Review of Ecology and Systematics 16: 393–430.

[WWN03] Wahlberg, N., E. Weingartner & S. Nylin. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 28: 473–484.

Last updated: 18 August 2017.

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