Common evening brown Melanitis leda, photographed by Ravi Vaidyanathan.

Belongs within: Nymphalidae.
Contains: Elymniini, Satyrini.

The Satyrinae is a cosmopolitan group of butterflies whose larvae have bifid tails and mostly feed on monocots. Most satyrines are relatively drab in coloration, with browns and greys predominating, though there are some more colourful tropical species.

<==Satyrinae [Satyridae]
    |--+--Elymniini WWN03
    |  `--Satyrini WWN03
    `--+--Manataria maculata WWN03
       `--Melanitis [Melanitini] WWN03
            |--M. constantia B88
            |--M. leda WWN03
            |    |--M. l. leda PP02
            |    |--M. l. levuna PP02
            |    `--M. l. saipanensis Nakamura 1929 TYM08
            `--M. undularis WM66

Satyrinae incertae sedis:
  Haetera GE05 [Haeterini WWN03]
    `--H. macleannania GE05
  Eritini WWN03
  Ragadiini WWN03
  Taygetis S55
    |--T. andromeda (Cramer 1779) S55
    |--T. kerea Butler 1869 FS54
    `--T. uncinata Weymer 1907 FS54
  Pierella lena (Linnaeus 1767) S55
  Bia [Biini] S55
    `--B. actorion (Linnaeus 1767) S55
  Dodonidia helmsi T27
  Neorinella garciae GE05
  Lethe GE05
    |--L. corbieri GE05
    `--L. diana I92
         |--L. d. diana I92
         `--L. d. mikuraensis Shirôzu 1975 I92
  Pseudoneorina coulleti GE05
  Tisiphone abeona NS04
    |--T. a. abeona NS04
    |--T. a. morrisi NS04
    |--T. a. rawnsleyi NS04
    `--T. a. regalis NS04
  Calisto L88
    |--C. hysius L88
    `--+--C. arcas L88
       `--C. archebates L88
  Eretris L88

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[FS54] Franz, E., & H. Schröder. 1954. Tagfalter (Lep. Rhopalocera) aus El Salvador. Senckenbergiana Biologica 35: 75-87.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385-409.

[NS04] New, T. R., & D. P. A. Sands. 2004. Management of threatened insect species in Australia, with particular reference to butterflies. Australian Journal of Entomology 43 (3): 258-270.

[PP02] Patrick, B., & H. Patrick. 2002. Butterflies of Fiji. Weta 24: 5-12.

[S55] Schröder, H. 1955. Eine Falter-Ausbeute aus dem westlichen Bolivien. (Ins. Lepid. Rhopal.) Senckenbergiana Biologica 36: 329-338.

[TYM08] Tennent, W. J., M. Yasuda & K. Morimoto. 2008. Lansania Journal of arachnology and zoology – a rare and obscure Japanese natural history journal. Archives of Natural History 35 (2): 252-280.

[T27] Thomson, G. M. 1927. The pollination of New Zealand flowers by birds and insects. Transactions and Proceedings of the New Zealand Institute 57: 106-125.

[WWN03] Wahlberg, N., E. Weingartner & S. Nylin. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 28: 473-484.

[WM66] Wallace, A. R., & F. Moore. 1866. List of lepidopterous insects collected at Takow, Formosa, by Mr. R. Swinhoe. Proceedings of the Zoological Society of London 1866: 355-365.

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