Lohora (Lohora) ophthalmicus, photographed by Antonio Guidici.

Belongs within: Satyrinae.

The Elymniini are a group of butterflies including the bushbrowns and palmflies.

    |  i. s.: Bicyclus anynana WWN03
    |         Lasiommata WWN03
    |           |--L. dejanira B66
    |           `--L. megera WWN03
    |         ‘Papilio’ canthus Linné 1766 (see below for synonymy) B66
    `--Mycalesina V-WF03
         |--Orsotriaena medus V-WF03
         |--Nirvana Tsukada & Nishiyama 1979 V-WF03
         |    `--N. hypnus V-WF03
         |--Mycalesis V-WF03
         |    |--*M. francisca [=Papilio francisca] V-WF03
         |    |--M. drusia [incl. M. mineus] WM66
         |    |--M. otrea WM66
         |    |--‘Papilio’ peribaea Fabricius 1793 [=Neonympha peribaea, Satyrus peribaea] B66
         |    `--M. samba [incl. M. lalassis] WM66
         `--Lohora Moore 1880 [=Lohara (l. c.); incl. Celebina Fruhstorfer 1899] V-WF03
              |--L. (Lohora) V-WF03
              |    |--*L. (L.) dexamenus (Hewitson 1862) [=Mycalesis dexamenus] V-WF03
              |    |--L. (L.) anna Vane-Wright & Fermon 2003 V-WF03
              |    |--L. (L.) dinon (Hewitson 1862) [=Mycalesis dinon, M. dexamenus dinon] V-WF03
              |    |--L. (L.) haasei (Röber 1887) [=Mycalesis haasei] V-WF03
              |    |--L. (L.) ophthalmicus (Westwood 1888) (see below for synonymy) V-WF03
              |    |--L. (L.) tilmara (Fruhstorfer 1906) [=Mycalesis tilmara] V-WF03
              |    |--L. (L.) transiens (Fruhstorfer 1908) (see below for synonymy) V-WF03
              |    `--L. (L.) unipapillata Fruhstorfer 1899 (see below for synonymy) V-WF03
              |--L. (Physcon de Nicéville 1898) [=Phycona (l. c.)] V-WF03
              |    |--L. (*P.) pandaea (Hopffer 1874) [=Mycalesis pandaea, M. (Physcon) pandaea] V-WF03
              |    |--L. (P.) decipiens (Martin 1929) [=Mycalesis decipiens] V-WF03
              |    |--L. (P.) deianira (Hewitson 1862) [=Mycalesis deianira, M. dora Hewitson 1864] V-WF03
              |    |--L. (P.) deianirina (Fruhstorfer 1897) (see below for synonymy) V-WF03
              |    |--L. (P.) erna (Fruhstorfer 1899) [=Celebina erna, Mycalesis erna] V-WF03
              |    |--L. (P.) imitatrix (Martin 1929) [=Mycalesis imitatrix] V-WF03
              |    |--L. (P.) inga (Fruhstorfer 1899) [=*Celebina inga, Mycalesis inga] V-WF03
              |    `--L. (P.) umbrosa Roos 1997 V-WF03
              `--L. (Pseudomycalesis Tsukada & Nishiyama 1979) V-WF03
                   `--L. (*P.) tanuki (Tsukada & Nishiyama 1979) [=*Pseudomycalesis tanuki] V-WF03

Lohora (Lohora) ophthalmicus (Westwood 1888) [=Messaras ophthalmicus, Mycalesis ophthalmica, My. ophthalmicus] V-WF03

Lohora (Lohora) transiens (Fruhstorfer 1908) [=Mycalesis dexamenus transiens; incl. M. transiens f. obscurata Martin 1929] V-WF03

Lohora (Lohora) unipapillata Fruhstorfer 1899 [=Lohara (l. c.) dinon unipapillata, Mycalesis haasei unipapillata] V-WF03

Lohora (Physcon) deianirina (Fruhstorfer 1897) [=Mycalesis deianirina, Celebina pandaea deianirina, Lohora pandaea deianirina, Mycalesis deianira deianirina] V-WF03

‘Papilio’ canthus Linné 1766 [=Neonympha canthus, Satyrus canthus, S. cantheus; incl. Papilio eurydice Linn. 1789] B66

*Type species of generic name indicated


[B66] Butler, A. G. 1866. A monograph of the genus Euptychia, a numerous race of butterflies belonging to the family Satyridae; with descriptions of sixty species new to science, and notes on their affinities &c. Proceedings of the Zoological Society of London 1866: 458-504.

[V-WF03] Vane-Wright, R. I., & H. Fermon. 2003. Taxonomy and identification of Lohora Moore (Lepidoptera: Satyrinae), the Sulawesi bush browns. Invertebrate Systematics 17: 129-141.

[WWN03] Wahlberg, N., E. Weingartner & S. Nylin. 2003. Towards a better understanding of the higher systematics of Nymphalidae (Lepidoptera: Papilionoidea). Molecular Phylogenetics and Evolution 28: 473-484.

[WM66] Wallace, A. R., & F. Moore. 1866. List of lepidopterous insects collected at Takow, Formosa, by Mr. R. Swinhoe. Proceedings of the Zoological Society of London 1866: 355-365.

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