Harpalinae

Calophaena cf. bicincta, copyright Udo Schmidt.


Belongs within: Conjunctae.
Contains: Agonocheila, Trigonothops, Coptocarpus, Darodilia, Leiradira, Trichosternus, Castelnaudia, Nurus, Sarticus, Prosopogmus, Simodontus, Rhytisternus, Morion, Zuphiitae, Abacetus, Loxandrini, Pterostichini, Calathus, Licinini, Harpalitae, Zabrini, Odacanthini, Pseudomorphini, Platynini, Oodini, Lachnophorini, Lebiini, Badister, Perigona.

The Harpalinae is a large clade of carabid ground beetles, distinguished by the distinctive forms of the parameres in the male genitalia. Over 19,000 species of Harpalinae are known from the modern fauna, exhibiting a wide range of ecologies including herbivorous and myrmecophilous forms. These species have been arranged in a correspondingly large number of tribes, but relationships between the tribes have been poorly known (Ober & Maddison 2008). These include the Morionini, a group of species with moniliform antennae, a projecting dentiform process above the antennal insertion, and usually protruding eyes. The Hexagoniini are a Palaeotropical group with a depressed body, a large head with cheek-like orbits and elongate mandibles, and pad-shaped tarsi. The Neotropical Ctenodactylini are found climbing on the stems of monocotyledons such as grasses and sedges, and possess adhesive setae on the underside of the tarsi (Ball & Bousquet 2001).

Synapomorphies (from Roig-Juñent & Cicchino 2001): Mandibular scrobal setae absent; maxillary palpomere 4 fusiform; elytral plica absent; left paramere conchiferous, differing in shape from right; right paramere glabrous.

<==Harpalinae (see below for synonymy)
    |--+--Moriosomus OM08
    |  `--Morionini [Morionides] B14
    |       |--Morion OM08
    |       |--Megamorio B14
    |       |--Hyperion Castelnau 1834 B14, LM87 (see below for synonymy)
    |       |    `--*H. schroetteri (Schreibers 1802) (see below for synonymy) LM87
    |       `--Platynodes westermanni Westwood 1847 B14
    |            |--P. w. westermanni B14
    |            `--P. w. peregrinus B14
    `--+--+--Zuphiitae OM08
       |  `--+--Hexagoniini [Hexagoniinae] OM08
       |     |    |--Dinopelma immaculatum OM08
       |     |    `--Hexagonia OM08
       |     |         |--H. angustula B55
       |     |         `--H. scabricollis B55
       |     `--Ctenodactylini [Ctenodactylitae] OM08
       |          |  i. s.: Calophaenoidea Liebke 1930 BB01
       |          |           `--C. arrowi B14
       |          |--Teukrus cruciatus OM08
       |          `--+--Ctenodactyla batesi OM08
       |             `--Leptotrachelus Latreille 1829 OM08, BB01 [incl. Spheracra Say 1830 BB01]
       |                  `--L. dorsalis OM08
       `--+--+--Metiini OM08
          |  |    |--Abropus carnifex OM08
          |  |    `--Metius OM08
          |  `--+--+--Abacetus OM08
          |     |  `--Loxandrini OM08
          |     `--+--+--Pterostichini OM08
          |        |  `--Cnemalobus [Cnemalobini] OM08
          |        |       `--C. sulciferus OM08
          |        `--+--Calathus OM08
          |           `--Synuchus Gyllenhal 1810 OM08, BB01
          |                |--S. dubius (LeConte 1854) BB01
          |                |--S. impunctatus (Say 1823) BB01
          |                `--S. semirufus Casey 1913 BB01
          `--+--+--Licinini OM08
             |  `--Harpalitae OM08
             `--+--Zabrini OM08
                `--+--+--Odacanthini OM08
                   |  `--Peleciini [Peleciinae] OM08
                   |       |--Pelecium sulcipenne OM08
                   |       |--Disphericus tarsalis B55
                   |       `--Eripus nitidus OM08
                   `--+--+--Pseudomorphini OM08
                      |  `--Platynini OM08
                      `--+--+--Oodini OM08
                         |  `--+--Lachnophorini OM08
                         |     `--Calophaena Klug 1821 OM08, BB01 [Calophaenini]
                         |          |--C. bicincta B14
                         |          |    |--C. b. bicincta B14
                         |          |    `--C. b. ligata Bates 1883 B14
                         |          `--C. dupuisi OM08
                         `--+--Lebiini OM08
                            `--+--+--Sinurus opacus OM08
                               |  `--Stenotelus opacus OM08
                               `--+--+--Cyrtolaus L88
                                  |  `--+--Barylaus L88
                                  |     `--Caelostomus Macleay 1825 OM08, LM87 (see below for synonymy)
                                  |          |--*C. picipes (Macleay 1825) (see below for synonymy) LM87
                                  |          |--C. albertisi Straneo 1938 LM87
                                  |          `--C. subsinuatus (Chaudoir 1883) [=Drimostoma subsinuata] LM87
                                  `--+--Badister OM08
                                     `--Perigonini [Perigoninae, Perigonitae, Trechicinae] OM08
                                          |  i. s.: Ripogenites Basilewsky 1954 BB01
                                          |--Perigona OM08
                                          `--+--Diploharpus laevissimus OM08
                                             `--Ripogena bembidioides [=Perigona (Ripogena) bembidioides] OM08

Harpalinae incertae sedis:
  Cillenum Samouelle 1819 B27
    |--C. chalmeri B27
    `--C. tillyardi Brookes 1927 B27
  Dichaetochilus Basilewsky 1946 B55
    |--D. incrassatus B55
    |--D. moestus B55
    `--D. planicollis B55
  Hyparpalus B55
    |--H. conviva B55
    `--H. holosericeus B55
  Laparhetes gracilis B55
  Platymetopus B55
    |--P. figuratus B55
    `--P. platythorax B55
  Neosiopelus Basilewsky 1946 B55
    `--N. punctatellus B55
  Aulacoryssus B55
    |--A. natalicus B55
    `--A. venustulus B55
  Xenodochus melanarius B55
    |--X. m. melanarius B55
    `--X. m. sissekabanus Basilewsky 1946 B55
  Ooidius dorsiger B55
  Bradybaenus Dejean 1829 B55, BR17 [Bradybaenina]
    `--B. opulentus B55
         |--B. o. opulentus B55
         `--B. o. insularis B55
  Microlestia B55
    |--M. atrata B55
    |--M. cribricollis B55
    `--M. immerita B55
  Netrodera formicaria B55
    |--N. f. formicaria B55
    |--N. f. damarensis B55
    `--N. f. peringueyi B55
  Thermophilum Basilewsky 1950 B55
    |--T. crudele B55
    |    |--T. c. crudele B55
    |    |--T. c. bennigseni B55
    |    `--T. c. westermanni B55
    |--T. homoplatum B55
    `--T. limbatum B55
         |--T. l. limbatum B55
         |--T. l. machadoi Basilewsky 1955 B55
         `--T. l. ovampoa B55
  Ginema Ball & Shpeley 2002 [Ginemini] BS02
    `--*G. thomasi Ball & Shpeley 2002 BS02
  Gestroania Liebke 1938 LM87
    `--*G. amplipennis (Gestro 1875) [=Casnonia amplipennis] LM87
  Myrmecodemus Sloane 1923 LM87
    |--*M. riverinae (Sloane 1890) [=Casnonia riverinae] LM87
    |--M. formicoides (Sloane 1910) [=Lachnothorax formicoides] LM87
    `--M. globulicollis (Macleay 1888) [=Casnonia globulicollis] LM87
  Anasis Castelnau 1867 LM87
    `--*A. howittii Castelnau 1867 LM87
  Aulacolius Sloane 1923 LM87
    `--*A. triordinatus Sloane 1923 LM87
  Deipyrus Liebke 1938 LM87
    `--*D. palustris (Sloane 1910) [=Lachnothorax palustris] LM87
  Basistichus Sloane 1917 LM87
    `--*B. micans (Macleay 1864) [=Odacantha micans] LM87
  Renneria kamouni GWW10
  Velindopsis panagaeoides B55
  Singiliomimus modestus B55
  Neocoptodera overlaeti B55
  Coptoderina B55
    |--C. amoenula B55
    |--C. congoana B55
    `--C. rugiceps B55
  Metalebia B55
    |--M. antelobata Basilewsky 1955 B55
    |--M. promontorii B55
    |--M. transvaalensis B55
    `--M. vaciva B55
  Coptoglossus Chaudoir 1869 LM87
    |--*C. sulcatulus Chaudoir 1869 LM87
    `--C. carteri (Sloane 1915) [=Platynus carteri] LM87
  Holcoderus Chaudoir 1869 LM87
    |--*H. praemorsus Chaudoir 1869 LM87
    `--H. caeruleipennis Sloane 1910 LM87
  Agonocheila LM87
  Minuthodes Andrewes 1941 [=Platia Chaudoir 1869 non HĂĽbner 1820] LM87
    |--*M. lineella (Chaudoir 1869) [=*Platia lineella] LM87
    `--M. queenslandica (Sloane 1917) [=Platia queenslandica] LM87
  Trigonothops LM87
  Speotarus Moore 1964 LM87
    `--*S. lucifugus Moore 1964 LM87
         |--S. l. lucifugus LM87
         `--S. l. princeps Moore 1964 LM87
  Aristolebia Bates 1892 LM87
    |--*A. quadridentata Bates 1892 LM87
    `--A. wau Darlington 1968 LM87
  Creagris Nietner 1857 [incl. Pseudhelluo Castelnau 1867] LM87
    |--*C. labrosa Nietner 1857 LM87
    `--C. wilsonii (Castelnau 1867) [=*Pseudhelluo wilsonii] LM87
  Helluonidius Chaudoir 1872 LM87
    |--*H. cyanipennis (Hope 1842) (see below for synonymy) LM87
    |--H. aterrimus (Macleay 1873) [=Helluosoma aterrimum] LM87
    |--H. cyaneus (Castelnau 1867) [=Helluosoma cyanea] LM87
    `--H. latipennis (Macleay 1887) [=Helluosoma latipenne] LM87
  Helluosoma Castelnau 1867 (see below for synonymy) LM87
    |--*H. atrum Castelnau 1867 [=H. ater] LM87
    `--H. longicolle Macleay 1888 LM87
  Helluodema Castelnau 1867 [incl. Simoglossus Chaudoir 1872] LM87
    |--H. unicolor (Hope 1842) (see below for synonymy) LM87
    `--H. brunneum Sloane 1917 LM87
  Epimicodema Sloane 1914 LM87
    `--*E. mastersii (Macleay 1871) [=Helluosoma mastersii] LM87
  Helluapterus Sloane 1914 LM87
    `--*H. niger Sloane 1914 [incl. H. niger punctata Sloane 1914] LM87
  Neohelluo Sloane 1914 LM87
    `--*N. angulicollis Sloane 1914 LM87
  Ametroglossus Sloane 1914 LM87
    `--*A. ater (Macleay 1887) [=Gigadema atrum] LM87
  Hololeius LaFertĂ©-SĂ©nectère 1851 [=Hololius Chaudoir 1857] LM87
    `--H. ceylanicus (Nietner 1856) (see below for synonymy) LM87
  Coptocarpus LM87
  Siagonyx Macleay 1871 LM87
    |--*S. amplipennis Macleay 1871 LM87
    |--S. blackburni Sloane 1916 LM87
    `--S. mastersii Macleay 1871 LM87
  Hormacrus Sloane 1898 LM87
    |--*H. latus Sloane 1898 LM87
    `--H. minor (Blackburn 1890) [=Lestignathus minor] LM87
  Platylytron Macleay 1873 [=Platelytron Csiki 1931] LM87
    `--*P. amplipenne Macleay 1873 LM87
  Dilonchus Andrewes 1936 LM87
    |--*D. bidens Andrewes 1936 LM87
    `--D. pictus Darlington 1963 LM87
  Omestes Andrewes 1933 LM87
    `--*O. torta Andrewes 1933 LM87
  Catadromus Macleay 1825 LM87
    |--*C. tenebrioides (Olivier 1790) [=Carabus tenebrioides; incl. Cat. elseyi White 1859] LM87
    |--C. australis Castelnau 1834 [incl. C. latro TschitschĂ©rine 1896] LM87
    |--C. goliath TschitschĂ©rine 1896 LM87
    |--C. lacordairei Boisduval 1835 LM87
    |--C. simonae Dupuis 1911 LM87
    `--C. tschitscherini Straneo 1941 LM87
  Cratogaster Blanchard 1853 (see below for synonymy) LM87
    |--*C. sulcata Blanchard 1853 [incl. Cyphosoma latum Chaudoir 1878] LM87
    |--C. melas (Castelnau 1867) (see below for synonymy) LM87
    |--C. occidentalis Macleay 1888 LM87
    |--C. robusta (Macleay 1883) [=Tibarisus robustus] LM87
    `--C. unicolor (Hope 1842) [=*Cyphosoma unicolor] LM87
  Liopasa TschitschĂ©rine 1901 LM87
    `--*L. crepera TschitschĂ©rine 1901 LM87
  Loxogenius Sloane 1907 LM87
    `--*L. opacipennis (Macleay 1887) (see below for synonymy) LM87
  Darodilia LM87
  Delinius Westwood 1864 LM87
    |--*D. essingtoni Westwood 1864 LM87
    `--D. castelnaui Chaudoir 1878 LM87
  Leiradira LM87
  Lesticus Dejean 1828 LM87
    |  i. s.: L. politus M86
    |--*L. (Lesticus) janthinus Dejean 1828 LM87
    `--L. (Triplogenius Chaudoir 1852) LM87
         |--‘Omasens’ viridicollis Macleay 1825 (see below for synonymy) LM87
         |--‘Triplogenius’ chalcothorax B89
         |--L. (T.) chloronotus Chaudoir 1868 LM87
         `--‘Triplogenius’ mouhotii B89
  Notolestus Sloane 1895 LM87
    `--*N. sulcipennis (Macleay 1873) [=Abax sulcipennis] LM87
  Megadromus Motschulsky 1865 [incl. Nesopterostichus TschitschĂ©rine 1902] LM87
    |  i. s.: ‘Platysma’ australasiae GuĂ©rin-MĂ©neville 1841 (see below for synonymy) LM87
    |         M. rectangulus WH02
    |         M. temukensis WH02
    |--M. (Megadromus) antarcticus (Chaudoir 1865) (see below for synonymy) LM87
    `--M. (Protodromus Moore 1965) LM87
         |--M. (*P.) eborensis Moore 1965 LM87
         `--M. (P.) australicus (Sloane 1916) [=Trichosternus australicus] LM87
  Trichosternus LM87
  Castelnaudia LM87
  Nurus LM87
  Paranurus TschitschĂ©rine 1901 LM87
    |--*P. dilaticeps (Chaudoir 1865) [=Feronia (Trichosternus) dilaticeps] LM87
    |--P. macleayi (Sloane 1895) [=Mecynognathus macleayi] LM87
    `--P. petri TschitschĂ©rine 1901 LM87
  Mecynognathus Macleay 1873 LM87
    `--*M. damelii Macleay 1873 LM87
  Notabax Moore 1976 LM87
    `--*N. monteithi Moore 1976 LM87
  Cratoferonia TschitschĂ©rine 1902 [=Ceratoferonia (l. c.)] LM87
    |--*C. regalis (Castelnau 1867) [=Feronia regalis] LM87
    `--C. phylarchus (Sloane 1900) [=Pterostichus phylarchus] LM87
  Secatophus Castelnau 1867 (see below for synonymy) LM87
    `--*S. australis (Hope 1845) (see below for synonymy) LM87
  Teropha Castelnau 1867 [incl. Morphnos Schaufuss 1867] LM87
    |--T. sturtii (White 1859) (see below for synonymy) LM87
    `--T. besti (Sloane 1902) [=Morphnos besti] LM87
  Acanthoferonia Moore 1965 LM87
    `--*A. ferox Moore 1965 LM87
  Sarticus LM87
  Rhabdotus Chaudoir 1865 LM87
    `--*R. reflexus (Chaudoir 1865) (see below for synonymy) LM87
  Loxodactylus Chaudoir 1865 LM87
    `--*L. carinatulus (Chaudoir 1865) [=Feronia (*Loxodactylus) carinatula] LM87
         |--L. c. carinatulus [incl. Feronia dingo Castelnau 1867] LM87
         `--L. c. amaeropterus (Chaudoir 1865) (see below for synonymy) LM87
  Conchitella Moore 1962 LM87
    `--*C. clivinoides Moore 1962 LM87
  Litarthrum Sloane 1915 LM87
    `--*L. browni Sloane 1915 LM87
  Gastrogmus Sloane 1915 LM87
    `--*G. ischialis Sloane 1915 LM87
  Prosopogmus LM87
  Simodontus LM87
  Setalis Castelnau 1867 [incl. Loxogmus Sloane 1890] LM87
    |--*S. niger Castelnau 1867 [incl. *Loxogmus obscurus Sloane 1890] LM87
    |--S. rubripes Sloane 1907 LM87
    `--S. sloanei Darlington 1953 LM87
  Setalimorphus Sloane 1895 LM87
    |--*S. punctiventris Sloane 1895 LM87
    `--S. regularis Sloane 1910 LM87
  Phaenaulax TschitschĂ©rine 1898 LM87
    `--P. nanus (Sloane 1895) [=Setalimorphus nanus; incl. *P. stenomorphus TschitschĂ©rine 1898] LM87
  Rhytisternus LM87
  Eurystomis Chaudoir 1878 LM87
    `--*E. castelnaui Chaudoir 1878 LM87
  Platycoelus Blanchard 1853 (see below for synonymy) LM87
    |--*P. depressus Blanchard 1853 [incl. *Dalbertisia lucidula Straneo 1939] LM87
    |--P. interstitialis (Sloane 1910) [=Chlaenioidius interstitialis] LM87
    |--*Hypherpinus’ jedlickai Straneo 1938 LM87
    |--P. melliei (Montrouzier 1860) (see below for synonymy) LM87
    |--P. poeciloides (Chaudoir 1878) (see below for synonymy) LM87
    `--P. prolixus (Erichson 1842) (see below for synonymy) LM87
  Nelidus Chaudoir 1878 LM87
    `--*N. australis Chaudoir 1878 LM87
  Zeodera Castelnau 1867 LM87
    `--*Z. atra Castelnau 1867 [=Z. ater] LM87
  Feronista Moore 1965 LM87
    |--*F. amaroides Moore 1965 LM87
    |--F. intermedia Moore 1965 LM87
    `--F. minor Moore 1965 LM87
  Pseudoceneus TschitschĂ©rine 1890 LM87
    |--P. (Pseudoceneus) [incl. Leptopodus Chaudoir 1874 non Schinz 1822] LM87
    |    |--*P. (P.) ‘holomelanus’ (TschitschĂ©rine 1890) [=Argutor holomelanus non Germar 1848] LM87
    |    `--P. (P.) sollicitus (Erichson 1842) (see below for synonymy) LM87
    `--P. (Poeciliridius Straneo 1937) LM87
         |--P. (*P.) iridescens (Castelnau 1867) [=Feronia iridescens] LM87
         `--P. (P.) norfolkensis Moore 1985 LM87
  Cuneipectus Sloane 1907 LM87
    |--*C. frenchi Sloane 1907 LM87
    `--C. foveatus Sloane 1915 LM87
  Geobaenus Dejean 1829 LM87
    |--*G. lateralis Dejean 1829 LM87
    `--G. australasiae GuĂ©rin-MĂ©neville 1839 LM87
  Cosmodiscus Sloane 1907 LM87
    `--*C. rubripictus Sloane 1907 LM87
  Brachidius Chaudoir 1852 LM87
    `--*B. crassicornis Chaudoir 1852 [incl. B. corpulentus Chaudoir 1872] LM87
  Fouquetius Maindron 1906 (see below for synonymy) LM87
    |--*F. rufus (Chaudoir 1876) [=*Holconotus (sensu Chaudoir) rufus] LM87
    |--*Holconotus’ (sensu Schmidt-Goebel) ferrugineus Schmidt-Goebel 1846 (n. o.) LM87
    `--F. variabilis Straneo 1960 LM87
  Pediomorphus Chaudoir 1878 LM87
    |--*P. planiusculus Chaudoir 1878 LM87
    |--‘Cyrtoderus’ australasiae Hope 1841 LM87
    |--‘Omalosoma’ carbonicolor Motschulsky 1865 LM87
    |--P. elongatus Sloane 1898 LM87
    |--P. macleayi Sloane 1900 LM87
    `--P. ruficollis Sloane 1900 LM87
  Violagonum Darlington 1956 LM87
    `--*V. violaceum (Chaudoir 1844) (see below for synonymy) LM87
  Lorostema Motschulsky 1864 [=Lorastemma Gemminger & Harold 1868] LM87
    |--*L. alutacea Motschulsky 1864 LM87
    `--L. bothriophora (Redtenbacher 1867) (see below for synonymy) LM87
  Notagonum Darlington 1952 LM87
    |--*N. externum Darlington 1952 LM87
    |--N. dentellum Darlington 1952 LM87
    |--N. lafertei (Montrouzier 1860) [=Anchomenus lafertei; incl. Platynus planipennis Macleay 1871] LM87
    |--N. macleayi (Sloane 1910) [=Platynus macleayi] LM87
    |--N. marginellum (Erichson 1842) (see below for synonymy) LM87
    |--N. murrayense (Blackburn 1890) [=Platynus murrayensis] LM87
    |--N. nigrellum Darlington 1956 LM87
    `--N. submetallicum (White 1846) (see below for synonymy) LM87
  Notoplatynus Moore 1985 LM87
    `--*N. darlingtoni Moore 1985 LM87
  Somoplatini BB01
    |--Somoplatus Dejean 1829 BB01
    |--Somoplatodes Basilewsky 1987 BB01
    |--Lophidius Dejean 1831 BB01
    `--Paralophidius Basilewsky 1987 BB01
  Percus paykullii LW89, B35
  ‘Ophionea’ Eschscholtz 1829 non Klug 1821 LM87
    |--*O. indica (Thunberg 1784) LM87, B14 [=Attelabus indica LM87]
    |--O. ishii OM08
    |--O. interstitialis B14
    `--O. nigrofasciata B14

Caelostomus Macleay 1825 OM08, LM87 [incl. Stomonaxus Motschulsky 1860 LM87; Caelostomini]

*Caelostomus picipes (Macleay 1825) [=Anaulacus (*Caelostomus) picipes; incl. Drimostoma rufipes Boheman 1858, Caelostomus striaticollis Chaudoir 1872] LM87

Cratogaster Blanchard 1853 [incl. Cyphosoma Hope 1842 nec Mannerheim 1837 nec Agassiz 1838, Pachidius Chaudoir 1865, Tibarisus Castelnau 1867] LM87

Cratogaster melas (Castelnau 1867) [=*Tibarisus melas; incl. T. ater Macleay 1871, Feronia (*Pachidius) sulcata Chaudoir 1865 non Cratogaster sulcata Blanchard 1853] LM87

Fouquetius Maindron 1906 [=Holconotus Chaudoir 1876 non Agassiz 1854; incl. Holconotus Schmidt-Goebel 1846 (n. o.) nec Agassiz 1854 nec Chaudoir 1876] LM87

Harpalinae [Callistitae, Chlaeniitae, Coptoderinae, Lebiinae, Lebiitae, Licinina, Masoreidae, Masoreomorphi, Odacanthinae, Odacanthitae, Pterostichinae, Pterostichitae]

Helluodema unicolor (Hope 1842) [=Aenigma unicolor; incl. Helluomorpha batesii Thomson 1857, *Helluodema batesii, Gigadema castelnaui Chaudoir 1872, *Simoglossus niger Chaudoir 1872] LM87

*Helluonidius cyanipennis (Hope 1842) [=Aenigma cyanipenne; incl. Helluosoma viridipenne Macleay 1887] LM87

Helluosoma Castelnau 1867 [incl. Helluodema Chaudoir 1872 non Castelnau 1867, Helluodoma (l. c.)] LM87

Hololeius ceylanicus (Nietner 1856) [=Chlaenius ceylanicus; incl. C. nitidulus Dejean 1826 non Schrank 1781, *Hololeius nitidulus, Hololius ornatus Tryon 1890] LM87

Hyperion Castelnau 1834 B14, LM87 [=Campylocnemis Westwood 1842 LM87, Heteroscelis Boisduval 1835 non Latreille 1829 LM87]

*Hyperion schroetteri (Schreibers 1802) [=Scarites schroetteri, *Campylocnemis schroetteri, *Heteroscelis schroetteri] LM87

Lorostema bothriophora (Redtenbacher 1867) [=Anchomenus (Agonum) bothriophora; incl. Platynus cooki Sloane 1895] LM87

Loxodactylus carinatulus amaeropterus (Chaudoir 1865) [=Feronia (Loxodactylus) amaeroptera; incl. F. yarrae Castelnau 1867] LM87

*Loxogenius opacipennis (Macleay 1887) [=Homalosoma opacipenne, Trichosternus (Castelnaudia) opacipennis] LM87

Megadromus (Megadromus) antarcticus (Chaudoir 1865) [=Feronia (Trichosternus) antarctica; incl. *M. viridilimbatus Motschulsky 1865] LM87

Notagonum marginellum (Erichson 1842) [=Anchomenus marginnellus; incl. Loxocrepis lugubris Motschulsky 1864, A. nigroaeneus Newman 1842] LM87

Notagonum submetallicum (White 1846) [=Colpodes submetallicum; incl. Platynus marginicollis Macleay 1871] LM87

‘Omasens’ viridicollis Macleay 1825 [incl. Trigonotoma bicolor Castelnau 1834, *Triplogenius bicolor] LM87

Platycoelus Blanchard 1853 [incl. Chlaenioidius Chaudoir 1865, Dalbertisia Straneo 1939, Hypherpinus Straneo 1938] LM87

Platycoelus melliei (Montrouzier 1860) [=Feronia melliei; incl. Poecilus chlaenioides Macleay 1888, F. (Chlaenioidius) herbacea Chaudoir 1865, F. resplendens Castelnau 1867 non Castelnau 1867] LM87

Platycoelus poeciloides (Chaudoir 1878) [=Feronia (Chlaenioidius) poeciloides; incl. F. (C.) irideomicans Tschitschérine 1890] LM87

Platycoelus prolixus (Erichson 1842) [=Pterostichus (Poecilus) prolixus, *Chlaenioidius prolixus; incl. Feronia funebris Castelnau 1867, C. planipennis Macleay 1871, Poecilus sulcatulus Macleay 1888] LM87

‘Platysma’ australasiae GuĂ©rin-MĂ©neville 1841 [incl. Platysma guerini Chaudoir 1865, Trichosternus (*Nesopterostichus) guerini] LM87

Pseudoceneus (Pseudoceneus) sollicitus (Erichson 1842) [=Pterostichus (Argutor) sollicitus; incl. Feronia interioris Castelnau 1867, F. iridipennis Castelnau 1867] LM87

*Rhabdotus reflexus (Chaudoir 1865) [=Feronia (*Rhabodotus) reflexa; incl. F. (R.) chaudoiri Tschitschérine 1890, F. diemenensis Castelnau 1867, Rhabdotus floridus Bates 1878] LM87

Secatophus Castelnau 1867 [incl. Hoploprion Tschitschérine 1903, Prionophorus Chaudoir 1865 non Blanchard 1853] LM87

*Secatophus australis (Hope 1845) [=Acinopus australis; incl. Feronia (*Prionophorus) crenatipes Chaudoir 1865, *Hoploprion crenatipes, Secatophus hopei Castelnau 1867] LM87

Teropha sturtii (White 1859) [=Platysma sturtii; incl. *Morphnos antipoda Schaufuss 1867, P. flindersii White 1859, *Teropha flindersii] LM87

*Violagonum violaceum (Chaudoir 1844) [=Harpalus violaceus, Colpodes violaceus; incl. C. mucronatus Macleay 1887] LM87

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[BS02] Ball, G. E., & D. Shpeley. 2002. Ginemini, Ginema thomasi, new tribe, new genus and new species, from Amazonian Bolivia (Coleoptera: Carabidae: Harpalinae). Transactions of the American Entomological Society 128 (1): 75–98.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B35] Boisduval, J. B. 1835. Voyage de DĂ©couvertes de l’Astrolabe. ExĂ©cutĂ© par ordre du Roi, pendant les annĂ©es 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'ocĂ©an Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. ColĂ©optères et autres ordres. J. Tastu: Paris.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. NĂĽtzel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B27] Brookes, A. E. 1927. A new genus and three new species of Coleoptera. Transactions and Proceedings of the New Zealand Institute 57: 563–566.

[GWW10] Guthrie, N. A., T. Weir & K. Will. 2010. Localised and regional patterns in ground-dwelling beetle assemblages in a semi-tropical arid zone environment. Records of the Western Australian Museum Supplement 78 (1): 169–184.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385–409.

[LW89] Lindquist, E. E., & D. E. Walter. 1989. Antennoseius (Vitzthumia) janus n.sp. (Acari: Ascidae), a mesostigmatic mite exhibiting adult female dimorphism. Canadian Journal of Zoology 67: 1291–1310.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

[WH02] Worthy, T. H., & R. N. Holdaway. 2002. The Lost World of the Moa: Prehistoric life of New Zealand. Indiana University Press: Bloomington (Indiana).

Last updated: 22 December 2020.

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