Harpalinae

Calophaena cf. bicincta, copyright Udo Schmidt.


Belongs within: Conjunctae.
Contains: Agonocheila, Trigonothops, Coptocarpus, Darodilia, Leiradira, Trichosternus, Castelnaudia, Nurus, Sarticus, Prosopogmus, Simodontus, Rhytisternus, Morion, Zuphiitae, Abacetus, Loxandrini, Pterostichini, Calathus, Licinini, Harpalitae, Zabrini, Odacanthini, Pseudomorphini, Platynini, Oodini, Lachnophorini, Lebiini, Badister, Perigona.

The Harpalinae is a large clade of carabid ground beetles, distinguished by the distinctive forms of the parameres in the male genitalia. Over 19,000 species of Harpalinae are known from the modern fauna, exhibiting a wide range of ecologies including herbivorous and myrmecophilous forms. These species have been arranged in a correspondingly large number of tribes, but relationships between the tribes have been poorly known (Ober & Maddison 2008). These include the Morionini, a group of species with moniliform antennae, a projecting dentiform process above the antennal insertion, and usually protruding eyes. The Hexagoniini are a Palaeotropical group with a depressed body, a large head with cheek-like orbits and elongate mandibles, and pad-shaped tarsi.

Synapomorphies (from Roig-Juñent & Cicchino 2001): Mandibular scrobal setae absent; maxillary palpomere 4 fusiform; elytral plica absent; left paramere conchiferous, differing in shape from right; right paramere glabrous.

<==Harpalinae (see below for synonymy)
    |--+--Moriosomus OM08
    |  `--Morionini [Morionides] B14
    |       |--Morion OM08
    |       |--Megamorio B14
    |       |--Hyperion Castelnau 1834 B14, LM87 (see below for synonymy)
    |       |    `--*H. schroetteri (Schreibers 1802) (see below for synonymy) LM87
    |       `--Platynodes westermanni Westwood 1847 B14
    |            |--P. w. westermanni B14
    |            `--P. w. peregrinus B14
    `--+--+--Zuphiitae OM08
       |  `--+--Hexagoniini [Hexagoniinae] OM08
       |     |    |--Dinopelma immaculatum OM08
       |     |    `--Hexagonia OM08
       |     |         |--H. angustula B55
       |     |         `--H. scabricollis B55
       |     `--Ctenodactylini [Ctenodactylitae] OM08
       |          |  i. s.: Calophaenoidea Liebke 1930 BB01
       |          |           `--C. arrowi B14
       |          |--Teukrus cruciatus OM08
       |          `--+--Ctenodactyla batesi OM08
       |             `--Leptotrachelus Latreille 1829 OM08, BB01 [incl. Spheracra Say 1830 BB01]
       |                  `--L. dorsalis OM08
       `--+--+--Metiini OM08
          |  |    |--Abropus carnifex OM08
          |  |    `--Metius OM08
          |  `--+--+--Abacetus OM08
          |     |  `--Loxandrini OM08
          |     `--+--+--Pterostichini OM08
          |        |  `--Cnemalobus [Cnemalobini] OM08
          |        |       `--C. sulciferus OM08
          |        `--+--Calathus OM08
          |           `--Synuchus Gyllenhal 1810 OM08, BB01
          |                |--S. dubius (LeConte 1854) BB01
          |                |--S. impunctatus (Say 1823) BB01
          |                `--S. semirufus Casey 1913 BB01
          `--+--+--Licinini OM08
             |  `--Harpalitae OM08
             `--+--Zabrini OM08
                `--+--+--Odacanthini OM08
                   |  `--Peleciini [Peleciinae] OM08
                   |       |--Pelecium sulcipenne OM08
                   |       |--Disphericus tarsalis B55
                   |       `--Eripus nitidus OM08
                   `--+--+--Pseudomorphini OM08
                      |  `--Platynini OM08
                      `--+--+--Oodini OM08
                         |  `--+--Lachnophorini OM08
                         |     `--Calophaena Klug 1821 OM08, BB01 [Calophaenini]
                         |          |--C. bicincta B14
                         |          |    |--C. b. bicincta B14
                         |          |    `--C. b. ligata Bates 1883 B14
                         |          `--C. dupuisi OM08
                         `--+--Lebiini OM08
                            `--+--+--Sinurus opacus OM08
                               |  `--Stenotelus opacus OM08
                               `--+--+--Cyrtolaus L88
                                  |  `--+--Barylaus L88
                                  |     `--Caelostomus Macleay 1825 OM08, LM87 (see below for synonymy)
                                  |          |--*C. picipes (Macleay 1825) (see below for synonymy) LM87
                                  |          |--C. albertisi Straneo 1938 LM87
                                  |          `--C. subsinuatus (Chaudoir 1883) [=Drimostoma subsinuata] LM87
                                  `--+--Badister OM08
                                     `--Perigonini [Perigoninae, Perigonitae, Trechicinae] OM08
                                          |  i. s.: Ripogenites Basilewsky 1954 BB01
                                          |--Perigona OM08
                                          `--+--Diploharpus laevissimus OM08
                                             `--Ripogena bembidioides [=Perigona (Ripogena) bembidioides] OM08

Harpalinae incertae sedis:
  Cillenum Samouelle 1819 B27
    |--C. chalmeri B27
    `--C. tillyardi Brookes 1927 B27
  Dichaetochilus Basilewsky 1946 B55
    |--D. incrassatus B55
    |--D. moestus B55
    `--D. planicollis B55
  Hyparpalus B55
    |--H. conviva B55
    `--H. holosericeus B55
  Laparhetes gracilis B55
  Platymetopus B55
    |--P. figuratus B55
    `--P. platythorax B55
  Neosiopelus Basilewsky 1946 B55
    `--N. punctatellus B55
  Aulacoryssus B55
    |--A. natalicus B55
    `--A. venustulus B55
  Xenodochus melanarius B55
    |--X. m. melanarius B55
    `--X. m. sissekabanus Basilewsky 1946 B55
  Ooidius dorsiger B55
  Bradybaenus Dejean 1829 B55, BR17 [Bradybaenina]
    `--B. opulentus B55
         |--B. o. opulentus B55
         `--B. o. insularis B55
  Microlestia B55
    |--M. atrata B55
    |--M. cribricollis B55
    `--M. immerita B55
  Netrodera formicaria B55
    |--N. f. formicaria B55
    |--N. f. damarensis B55
    `--N. f. peringueyi B55
  Thermophilum Basilewsky 1950 B55
    |--T. crudele B55
    |    |--T. c. crudele B55
    |    |--T. c. bennigseni B55
    |    `--T. c. westermanni B55
    |--T. homoplatum B55
    `--T. limbatum B55
         |--T. l. limbatum B55
         |--T. l. machadoi Basilewsky 1955 B55
         `--T. l. ovampoa B55
  Ginema Ball & Shpeley 2002 [Ginemini] BS02
    `--*G. thomasi Ball & Shpeley 2002 BS02
  Gestroania Liebke 1938 LM87
    `--*G. amplipennis (Gestro 1875) [=Casnonia amplipennis] LM87
  Myrmecodemus Sloane 1923 LM87
    |--*M. riverinae (Sloane 1890) [=Casnonia riverinae] LM87
    |--M. formicoides (Sloane 1910) [=Lachnothorax formicoides] LM87
    `--M. globulicollis (Macleay 1888) [=Casnonia globulicollis] LM87
  Anasis Castelnau 1867 LM87
    `--*A. howittii Castelnau 1867 LM87
  Aulacolius Sloane 1923 LM87
    `--*A. triordinatus Sloane 1923 LM87
  Deipyrus Liebke 1938 LM87
    `--*D. palustris (Sloane 1910) [=Lachnothorax palustris] LM87
  Basistichus Sloane 1917 LM87
    `--*B. micans (Macleay 1864) [=Odacantha micans] LM87
  Renneria kamouni GWW10
  Velindopsis panagaeoides B55
  Singiliomimus modestus B55
  Neocoptodera overlaeti B55
  Coptoderina B55
    |--C. amoenula B55
    |--C. congoana B55
    `--C. rugiceps B55
  Metalebia B55
    |--M. antelobata Basilewsky 1955 B55
    |--M. promontorii B55
    |--M. transvaalensis B55
    `--M. vaciva B55
  Coptoglossus Chaudoir 1869 LM87
    |--*C. sulcatulus Chaudoir 1869 LM87
    `--C. carteri (Sloane 1915) [=Platynus carteri] LM87
  Holcoderus Chaudoir 1869 LM87
    |--*H. praemorsus Chaudoir 1869 LM87
    `--H. caeruleipennis Sloane 1910 LM87
  Agonocheila LM87
  Minuthodes Andrewes 1941 [=Platia Chaudoir 1869 non Hübner 1820] LM87
    |--*M. lineella (Chaudoir 1869) [=*Platia lineella] LM87
    `--M. queenslandica (Sloane 1917) [=Platia queenslandica] LM87
  Trigonothops LM87
  Speotarus Moore 1964 LM87
    `--*S. lucifugus Moore 1964 LM87
         |--S. l. lucifugus LM87
         `--S. l. princeps Moore 1964 LM87
  Aristolebia Bates 1892 LM87
    |--*A. quadridentata Bates 1892 LM87
    `--A. wau Darlington 1968 LM87
  Creagris Nietner 1857 [incl. Pseudhelluo Castelnau 1867] LM87
    |--*C. labrosa Nietner 1857 LM87
    `--C. wilsonii (Castelnau 1867) [=*Pseudhelluo wilsonii] LM87
  Helluonidius Chaudoir 1872 LM87
    |--*H. cyanipennis (Hope 1842) (see below for synonymy) LM87
    |--H. aterrimus (Macleay 1873) [=Helluosoma aterrimum] LM87
    |--H. cyaneus (Castelnau 1867) [=Helluosoma cyanea] LM87
    `--H. latipennis (Macleay 1887) [=Helluosoma latipenne] LM87
  Helluosoma Castelnau 1867 (see below for synonymy) LM87
    |--*H. atrum Castelnau 1867 [=H. ater] LM87
    `--H. longicolle Macleay 1888 LM87
  Helluodema Castelnau 1867 [incl. Simoglossus Chaudoir 1872] LM87
    |--H. unicolor (Hope 1842) (see below for synonymy) LM87
    `--H. brunneum Sloane 1917 LM87
  Epimicodema Sloane 1914 LM87
    `--*E. mastersii (Macleay 1871) [=Helluosoma mastersii] LM87
  Helluapterus Sloane 1914 LM87
    `--*H. niger Sloane 1914 [incl. H. niger punctata Sloane 1914] LM87
  Neohelluo Sloane 1914 LM87
    `--*N. angulicollis Sloane 1914 LM87
  Ametroglossus Sloane 1914 LM87
    `--*A. ater (Macleay 1887) [=Gigadema atrum] LM87
  Hololeius LaFerté-Sénectère 1851 [=Hololius Chaudoir 1857] LM87
    `--H. ceylanicus (Nietner 1856) (see below for synonymy) LM87
  Coptocarpus LM87
  Siagonyx Macleay 1871 LM87
    |--*S. amplipennis Macleay 1871 LM87
    |--S. blackburni Sloane 1916 LM87
    `--S. mastersii Macleay 1871 LM87
  Hormacrus Sloane 1898 LM87
    |--*H. latus Sloane 1898 LM87
    `--H. minor (Blackburn 1890) [=Lestignathus minor] LM87
  Platylytron Macleay 1873 [=Platelytron Csiki 1931] LM87
    `--*P. amplipenne Macleay 1873 LM87
  Dilonchus Andrewes 1936 LM87
    |--*D. bidens Andrewes 1936 LM87
    `--D. pictus Darlington 1963 LM87
  Omestes Andrewes 1933 LM87
    `--*O. torta Andrewes 1933 LM87
  Catadromus Macleay 1825 LM87
    |--*C. tenebrioides (Olivier 1790) [=Carabus tenebrioides; incl. Cat. elseyi White 1859] LM87
    |--C. australis Castelnau 1834 [incl. C. latro Tschitschérine 1896] LM87
    |--C. goliath Tschitschérine 1896 LM87
    |--C. lacordairei Boisduval 1835 LM87
    |--C. simonae Dupuis 1911 LM87
    `--C. tschitscherini Straneo 1941 LM87
  Cratogaster Blanchard 1853 (see below for synonymy) LM87
    |--*C. sulcata Blanchard 1853 [incl. Cyphosoma latum Chaudoir 1878] LM87
    |--C. melas (Castelnau 1867) (see below for synonymy) LM87
    |--C. occidentalis Macleay 1888 LM87
    |--C. robusta (Macleay 1883) [=Tibarisus robustus] LM87
    `--C. unicolor (Hope 1842) [=*Cyphosoma unicolor] LM87
  Liopasa Tschitschérine 1901 LM87
    `--*L. crepera Tschitschérine 1901 LM87
  Loxogenius Sloane 1907 LM87
    `--*L. opacipennis (Macleay 1887) (see below for synonymy) LM87
  Darodilia LM87
  Delinius Westwood 1864 LM87
    |--*D. essingtoni Westwood 1864 LM87
    `--D. castelnaui Chaudoir 1878 LM87
  Leiradira LM87
  Lesticus Dejean 1828 LM87
    |  i. s.: L. politus M86
    |--*L. (Lesticus) janthinus Dejean 1828 LM87
    `--L. (Triplogenius Chaudoir 1852) LM87
         |--‘Omasens’ viridicollis Macleay 1825 (see below for synonymy) LM87
         |--‘Triplogenius’ chalcothorax B89
         |--L. (T.) chloronotus Chaudoir 1868 LM87
         `--‘Triplogenius’ mouhotii B89
  Notolestus Sloane 1895 LM87
    `--*N. sulcipennis (Macleay 1873) [=Abax sulcipennis] LM87
  Megadromus Motschulsky 1865 [incl. Nesopterostichus Tschitschérine 1902] LM87
    |  i. s.: ‘Platysma’ australasiae Guérin-Méneville 1841 (see below for synonymy) LM87
    |         M. rectangulus WH02
    |         M. temukensis WH02
    |--M. (Megadromus) antarcticus (Chaudoir 1865) (see below for synonymy) LM87
    `--M. (Protodromus Moore 1965) LM87
         |--M. (*P.) eborensis Moore 1965 LM87
         `--M. (P.) australicus (Sloane 1916) [=Trichosternus australicus] LM87
  Trichosternus LM87
  Castelnaudia LM87
  Nurus LM87
  Paranurus Tschitschérine 1901 LM87
    |--*P. dilaticeps (Chaudoir 1865) [=Feronia (Trichosternus) dilaticeps] LM87
    |--P. macleayi (Sloane 1895) [=Mecynognathus macleayi] LM87
    `--P. petri Tschitschérine 1901 LM87
  Mecynognathus Macleay 1873 LM87
    `--*M. damelii Macleay 1873 LM87
  Notabax Moore 1976 LM87
    `--*N. monteithi Moore 1976 LM87
  Cratoferonia Tschitschérine 1902 [=Ceratoferonia (l. c.)] LM87
    |--*C. regalis (Castelnau 1867) [=Feronia regalis] LM87
    `--C. phylarchus (Sloane 1900) [=Pterostichus phylarchus] LM87
  Secatophus Castelnau 1867 (see below for synonymy) LM87
    `--*S. australis (Hope 1845) (see below for synonymy) LM87
  Teropha Castelnau 1867 [incl. Morphnos Schaufuss 1867] LM87
    |--T. sturtii (White 1859) (see below for synonymy) LM87
    `--T. besti (Sloane 1902) [=Morphnos besti] LM87
  Acanthoferonia Moore 1965 LM87
    `--*A. ferox Moore 1965 LM87
  Sarticus LM87
  Rhabdotus Chaudoir 1865 LM87
    `--*R. reflexus (Chaudoir 1865) (see below for synonymy) LM87
  Loxodactylus Chaudoir 1865 LM87
    `--*L. carinatulus (Chaudoir 1865) [=Feronia (*Loxodactylus) carinatula] LM87
         |--L. c. carinatulus [incl. Feronia dingo Castelnau 1867] LM87
         `--L. c. amaeropterus (Chaudoir 1865) (see below for synonymy) LM87
  Conchitella Moore 1962 LM87
    `--*C. clivinoides Moore 1962 LM87
  Litarthrum Sloane 1915 LM87
    `--*L. browni Sloane 1915 LM87
  Gastrogmus Sloane 1915 LM87
    `--*G. ischialis Sloane 1915 LM87
  Prosopogmus LM87
  Simodontus LM87
  Setalis Castelnau 1867 [incl. Loxogmus Sloane 1890] LM87
    |--*S. niger Castelnau 1867 [incl. *Loxogmus obscurus Sloane 1890] LM87
    |--S. rubripes Sloane 1907 LM87
    `--S. sloanei Darlington 1953 LM87
  Setalimorphus Sloane 1895 LM87
    |--*S. punctiventris Sloane 1895 LM87
    `--S. regularis Sloane 1910 LM87
  Phaenaulax Tschitschérine 1898 LM87
    `--P. nanus (Sloane 1895) [=Setalimorphus nanus; incl. *P. stenomorphus Tschitschérine 1898] LM87
  Rhytisternus LM87
  Eurystomis Chaudoir 1878 LM87
    `--*E. castelnaui Chaudoir 1878 LM87
  Platycoelus Blanchard 1853 (see below for synonymy) LM87
    |--*P. depressus Blanchard 1853 [incl. *Dalbertisia lucidula Straneo 1939] LM87
    |--P. interstitialis (Sloane 1910) [=Chlaenioidius interstitialis] LM87
    |--*Hypherpinus’ jedlickai Straneo 1938 LM87
    |--P. melliei (Montrouzier 1860) (see below for synonymy) LM87
    |--P. poeciloides (Chaudoir 1878) (see below for synonymy) LM87
    `--P. prolixus (Erichson 1842) (see below for synonymy) LM87
  Nelidus Chaudoir 1878 LM87
    `--*N. australis Chaudoir 1878 LM87
  Zeodera Castelnau 1867 LM87
    `--*Z. atra Castelnau 1867 [=Z. ater] LM87
  Feronista Moore 1965 LM87
    |--*F. amaroides Moore 1965 LM87
    |--F. intermedia Moore 1965 LM87
    `--F. minor Moore 1965 LM87
  Pseudoceneus Tschitschérine 1890 LM87
    |--P. (Pseudoceneus) [incl. Leptopodus Chaudoir 1874 non Schinz 1822] LM87
    |    |--*P. (P.) ‘holomelanus’ (Tschitschérine 1890) [=Argutor holomelanus non Germar 1848] LM87
    |    `--P. (P.) sollicitus (Erichson 1842) (see below for synonymy) LM87
    `--P. (Poeciliridius Straneo 1937) LM87
         |--P. (*P.) iridescens (Castelnau 1867) [=Feronia iridescens] LM87
         `--P. (P.) norfolkensis Moore 1985 LM87
  Cuneipectus Sloane 1907 LM87
    |--*C. frenchi Sloane 1907 LM87
    `--C. foveatus Sloane 1915 LM87
  Geobaenus Dejean 1829 LM87
    |--*G. lateralis Dejean 1829 LM87
    `--G. australasiae Guérin-Méneville 1839 LM87
  Cosmodiscus Sloane 1907 LM87
    `--*C. rubripictus Sloane 1907 LM87
  Brachidius Chaudoir 1852 LM87
    `--*B. crassicornis Chaudoir 1852 [incl. B. corpulentus Chaudoir 1872] LM87
  Fouquetius Maindron 1906 (see below for synonymy) LM87
    |--*F. rufus (Chaudoir 1876) [=*Holconotus (sensu Chaudoir) rufus] LM87
    |--*Holconotus’ (sensu Schmidt-Goebel) ferrugineus Schmidt-Goebel 1846 (n. o.) LM87
    `--F. variabilis Straneo 1960 LM87
  Pediomorphus Chaudoir 1878 LM87
    |--*P. planiusculus Chaudoir 1878 LM87
    |--‘Cyrtoderus’ australasiae Hope 1841 LM87
    |--‘Omalosoma’ carbonicolor Motschulsky 1865 LM87
    |--P. elongatus Sloane 1898 LM87
    |--P. macleayi Sloane 1900 LM87
    `--P. ruficollis Sloane 1900 LM87
  Violagonum Darlington 1956 LM87
    `--*V. violaceum (Chaudoir 1844) (see below for synonymy) LM87
  Lorostema Motschulsky 1864 [=Lorastemma Gemminger & Harold 1868] LM87
    |--*L. alutacea Motschulsky 1864 LM87
    `--L. bothriophora (Redtenbacher 1867) (see below for synonymy) LM87
  Notagonum Darlington 1952 LM87
    |--*N. externum Darlington 1952 LM87
    |--N. dentellum Darlington 1952 LM87
    |--N. lafertei (Montrouzier 1860) [=Anchomenus lafertei; incl. Platynus planipennis Macleay 1871] LM87
    |--N. macleayi (Sloane 1910) [=Platynus macleayi] LM87
    |--N. marginellum (Erichson 1842) (see below for synonymy) LM87
    |--N. murrayense (Blackburn 1890) [=Platynus murrayensis] LM87
    |--N. nigrellum Darlington 1956 LM87
    `--N. submetallicum (White 1846) (see below for synonymy) LM87
  Notoplatynus Moore 1985 LM87
    `--*N. darlingtoni Moore 1985 LM87
  Somoplatini BB01
    |--Somoplatus Dejean 1829 BB01
    |--Somoplatodes Basilewsky 1987 BB01
    |--Lophidius Dejean 1831 BB01
    `--Paralophidius Basilewsky 1987 BB01
  Percus paykullii LW89, B35
  ‘Ophionea’ Eschscholtz 1829 non Klug 1821 LM87
    |--*O. indica (Thunberg 1784) LM87, B14 [=Attelabus indica LM87]
    |--O. ishii OM08
    |--O. interstitialis B14
    `--O. nigrofasciata B14

Caelostomus Macleay 1825 OM08, LM87 [incl. Stomonaxus Motschulsky 1860 LM87; Caelostomini]

*Caelostomus picipes (Macleay 1825) [=Anaulacus (*Caelostomus) picipes; incl. Drimostoma rufipes Boheman 1858, Caelostomus striaticollis Chaudoir 1872] LM87

Cratogaster Blanchard 1853 [incl. Cyphosoma Hope 1842 nec Mannerheim 1837 nec Agassiz 1838, Pachidius Chaudoir 1865, Tibarisus Castelnau 1867] LM87

Cratogaster melas (Castelnau 1867) [=*Tibarisus melas; incl. T. ater Macleay 1871, Feronia (*Pachidius) sulcata Chaudoir 1865 non Cratogaster sulcata Blanchard 1853] LM87

Fouquetius Maindron 1906 [=Holconotus Chaudoir 1876 non Agassiz 1854; incl. Holconotus Schmidt-Goebel 1846 (n. o.) nec Agassiz 1854 nec Chaudoir 1876] LM87

Harpalinae [Callistitae, Chlaeniitae, Coptoderinae, Lebiinae, Lebiitae, Licinina, Masoreidae, Masoreomorphi, Odacanthinae, Odacanthitae, Pterostichinae, Pterostichitae]

Helluodema unicolor (Hope 1842) [=Aenigma unicolor; incl. Helluomorpha batesii Thomson 1857, *Helluodema batesii, Gigadema castelnaui Chaudoir 1872, *Simoglossus niger Chaudoir 1872] LM87

*Helluonidius cyanipennis (Hope 1842) [=Aenigma cyanipenne; incl. Helluosoma viridipenne Macleay 1887] LM87

Helluosoma Castelnau 1867 [incl. Helluodema Chaudoir 1872 non Castelnau 1867, Helluodoma (l. c.)] LM87

Hololeius ceylanicus (Nietner 1856) [=Chlaenius ceylanicus; incl. C. nitidulus Dejean 1826 non Schrank 1781, *Hololeius nitidulus, Hololius ornatus Tryon 1890] LM87

Hyperion Castelnau 1834 B14, LM87 [=Campylocnemis Westwood 1842 LM87, Heteroscelis Boisduval 1835 non Latreille 1829 LM87]

*Hyperion schroetteri (Schreibers 1802) [=Scarites schroetteri, *Campylocnemis schroetteri, *Heteroscelis schroetteri] LM87

Lorostema bothriophora (Redtenbacher 1867) [=Anchomenus (Agonum) bothriophora; incl. Platynus cooki Sloane 1895] LM87

Loxodactylus carinatulus amaeropterus (Chaudoir 1865) [=Feronia (Loxodactylus) amaeroptera; incl. F. yarrae Castelnau 1867] LM87

*Loxogenius opacipennis (Macleay 1887) [=Homalosoma opacipenne, Trichosternus (Castelnaudia) opacipennis] LM87

Megadromus (Megadromus) antarcticus (Chaudoir 1865) [=Feronia (Trichosternus) antarctica; incl. *M. viridilimbatus Motschulsky 1865] LM87

Notagonum marginellum (Erichson 1842) [=Anchomenus marginnellus; incl. Loxocrepis lugubris Motschulsky 1864, A. nigroaeneus Newman 1842] LM87

Notagonum submetallicum (White 1846) [=Colpodes submetallicum; incl. Platynus marginicollis Macleay 1871] LM87

‘Omasens’ viridicollis Macleay 1825 [incl. Trigonotoma bicolor Castelnau 1834, *Triplogenius bicolor] LM87

Platycoelus Blanchard 1853 [incl. Chlaenioidius Chaudoir 1865, Dalbertisia Straneo 1939, Hypherpinus Straneo 1938] LM87

Platycoelus melliei (Montrouzier 1860) [=Feronia melliei; incl. Poecilus chlaenioides Macleay 1888, F. (Chlaenioidius) herbacea Chaudoir 1865, F. resplendens Castelnau 1867 non Castelnau 1867] LM87

Platycoelus poeciloides (Chaudoir 1878) [=Feronia (Chlaenioidius) poeciloides; incl. F. (C.) irideomicans Tschitschérine 1890] LM87

Platycoelus prolixus (Erichson 1842) [=Pterostichus (Poecilus) prolixus, *Chlaenioidius prolixus; incl. Feronia funebris Castelnau 1867, C. planipennis Macleay 1871, Poecilus sulcatulus Macleay 1888] LM87

‘Platysma’ australasiae Guérin-Méneville 1841 [incl. Platysma guerini Chaudoir 1865, Trichosternus (*Nesopterostichus) guerini] LM87

Pseudoceneus (Pseudoceneus) sollicitus (Erichson 1842) [=Pterostichus (Argutor) sollicitus; incl. Feronia interioris Castelnau 1867, F. iridipennis Castelnau 1867] LM87

*Rhabdotus reflexus (Chaudoir 1865) [=Feronia (*Rhabodotus) reflexa; incl. F. (R.) chaudoiri Tschitschérine 1890, F. diemenensis Castelnau 1867, Rhabdotus floridus Bates 1878] LM87

Secatophus Castelnau 1867 [incl. Hoploprion Tschitschérine 1903, Prionophorus Chaudoir 1865 non Blanchard 1853] LM87

*Secatophus australis (Hope 1845) [=Acinopus australis; incl. Feronia (*Prionophorus) crenatipes Chaudoir 1865, *Hoploprion crenatipes, Secatophus hopei Castelnau 1867] LM87

Teropha sturtii (White 1859) [=Platysma sturtii; incl. *Morphnos antipoda Schaufuss 1867, P. flindersii White 1859, *Teropha flindersii] LM87

*Violagonum violaceum (Chaudoir 1844) [=Harpalus violaceus, Colpodes violaceus; incl. C. mucronatus Macleay 1887] LM87

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[BS02] Ball, G. E., & D. Shpeley. 2002. Ginemini, Ginema thomasi, new tribe, new genus and new species, from Amazonian Bolivia (Coleoptera: Carabidae: Harpalinae). Transactions of the American Entomological Society 128 (1): 75–98.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B27] Brookes, A. E. 1927. A new genus and three new species of Coleoptera. Transactions and Proceedings of the New Zealand Institute 57: 563–566.

[GWW10] Guthrie, N. A., T. Weir & K. Will. 2010. Localised and regional patterns in ground-dwelling beetle assemblages in a semi-tropical arid zone environment. Records of the Western Australian Museum Supplement 78 (1): 169–184.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385–409.

[LW89] Lindquist, E. E., & D. E. Walter. 1989. Antennoseius (Vitzthumia) janus n.sp. (Acari: Ascidae), a mesostigmatic mite exhibiting adult female dimorphism. Canadian Journal of Zoology 67: 1291–1310.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

[WH02] Worthy, T. H., & R. N. Holdaway. 2002. The Lost World of the Moa: Prehistoric life of New Zealand. Indiana University Press: Bloomington (Indiana).

Last updated: 3 November 2018.

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