Caelostomus picipes, copyright Roman Dudko.

Belongs within: Harpalinae.
Contains: Notonomus, Abacetus.

The Pterostichini are a group of mostly mid-sized, relatively stout ground beetles, with antennae that are setose from the fourth segment onwards.

Pterostichini B90
    |--Cyrtolaus L88
    `--+--Barylaus L88
       `--Caelostomus Macleay 1825 LM87 [incl. Stomonaxus Motschulsky 1860 LM87; Caelostomi L88]
            |--*C. picipes (Macleay 1825) (see below for synonymy) LM87
            |--C. albertisi Straneo 1938 LM87
            `--C. subsinuatus (Chaudoir 1883) [=Drimostoma subsinuata] LM87

Pterostichini incertae sedis:
  Pterostichus B90
    |--P. anthracinus C01
    |--P. chalcites [=Poecilus chalcites] BB68
    |--P. cupreus (Linnaeus 1758) VPB11
    |--P. dilutipes C01
    |--P. eschscholtzii C01
    |--P. honestus B90
    |--P. madidus (Fabricius 1775) VPB11
    |--P. melanarius (Illiger 1798) VPB11
    |--P. metallicus FFB88
    |--P. (Piesmus) monedulus B90
    |--P. niger C01
    |--P. nigrita (Paykull 1790) VPB11
    |--P. oblongopunctatus C01
    |--P. pumilio FFB88
    |--P. vitreus C01
    `--P. vulgaris C01
  Synuchus B90
  Calathus B90
    |--C. ambiguus [incl. C. ambiguus var. dilutus] C01
    `--C. opacus Lucas 1846 E12
  Myas B90
  Oxycrepis ulkei B90
  Evarthrus B90
    |--E. gravesi B90
    `--E. sodalis BB68
  Abacidus B90
  Olisthopus B90
  Atranus pubescens B90
  Notonomus LB91
  Abacetus LB91

*Caeolostomus picipes (Macleay 1825) [=Anaulacus (*Caelostomus) picipes; incl. Drimostoma rufipes Boheman 1858, Caelostomus striaticollis Chaudoir 1872] LM87

*Type species of generic name indicated


[BB68] Basford, N. L., J. E. Butler, C. A. Leone & F. J. Rohlf. 1968. Immunologic comparisons of selected Coleoptera with analyses of relationships using numerical taxonomic methods. Systematic Zoology 17 (4): 388–406.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[FFB88] Franke, U., B. Friebe & L. Beck. 1988. Methodisches zur Ermittlung der Siedlungsdichte von Bodentieren aus Quadratproben und Barberfallen. Pedobiologia 32: 253–264.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385–409.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

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