Licinini

Dicaelus purpuratus, copyright Betsy Betros.

Belongs within: Harpalinae.
Contains: Lacordairia.

The Licinini are a cosmopolitan group of ground beetles characterised by a deep notch on the labrum.

<==Licinini [Licininae] M80
    |--+--Licinus Fabricius 1802 OM08, BR05
    |  |    |--L. cassideus [=Carabus cassideus] L02
    |  |    `--L. punctatulus [=Carabus punctatulus] L02
    |  `--Dicaelina OM08
    |       |--Diplocheila striatopunctata OM08
    |       `--Dicaelus OM08
    |            |--D. ambiguus OM08
    |            `--D. purpuratus Bonelli 1813 B14
    |                 |--D. p. purpuratus B14
    |                 `--D. p. splendidus B14
    `--+--Dicrochile Guérin-Méneville 1846 OM08, LM87 (see below for synonymy)
       |    |--*D. goryi (Boisduval 1835) (see below for synonymy) LM87
       |    |--D. brevicollis Chaudoir 1852 [=Dicronochilus brevicollis] LM87
       |    |--D. gigas Castelnau 1867 LM87
       |    |--D. idae Moore 1985 LM87
       |    |--D. minuta Castelnau 1867 LM87
       |    |--D. punctatostriata Castelnau 1867 LM87
       |    |--D. punctipennis Castelnau 1867 LM87
       |    |--D. punctulata Sloane 1923 LM87
       |    |--D. quadricollis Castelnau 1867 LM87
       |    `--D. ventralis Blackburn 1891 LM87
       `--+--Eutogeneius fuscus OM08
          `--Lestignathina OM08
               |--Lacordairia OM08
               `--Lestignathus Erichson 1842 LM87
                    |--*L. cursor Erichson 1842 LM87
                    |--L. foveatus Sloane 1920 LM87
                    `--L. simsoni Bates 1878 LM87

Licinini incertae sedis:
  Microferonia Blackburn 1890 M80, LM87
    |--*M. adelaidae Blackburn 1890 LM87
    |--M. anchomenoides (Macleay 1871) [=Badister anchomenoides, M. macleayi Csiki 1931] LM87
    |--M. cinctipennis Sloane 1898 LM87
    `--M. marginata (Castelnau 1867) [=Lacordairia marginata] LM87
  Physoloesthus Chaudoir 1850 M80, LM87 [=Physolaesthus LM87]
    |--*P. australis Chaudoir 1850 [=Physolaesthus australis] LM87
    |--P. caviceps (Andrewes 1936) [=Badister caviceps] LM87
    |--P. grandipalpis Macleay 1871 LM87
    |--P. insularis Bates 1878 [=Physolaesthus insularis] LM87
    |--P. pallidus Blackburn 1890 [=Physolesthus pallidus] LM87
    |--P. ruficollis Sloane 1900 [=Physolesthus ruficollis] LM87
    `--P. suturalis Castelnau 1867 [=Physoloestus suturalis] LM87

Dicrochile Guérin-Méneville 1846 OM08, LM87 [=Dicronochilus (l. c.) LM87; incl. Stomatocoelus Macleay 1864 LM87; Dicrochilina]

*Dicrochile goryi (Boisduval 1835) [=Rembus goryi; incl. *Stomatocoelus licinoides Macleay 1864, Dicrochile montana Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M80] Matthews, E. G. 1980. A Guide to the Genera of Beetles of South Australia. Part 1. Archostemata and Adephaga. South Australian Museum: Australia.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.