Agabinae

Agabus unguicularis, photographed by N. Sloth.

Belongs within: Dytiscidae.
Contains: Ilybius.

The Agabinae is a group of diving beetles sharing the characters of an externally visible scutellum, and metatarsal claws that are equal in length. Members of the Agabini are also united by the presence of a linear series of setae at the anteroventral angle of the metafemur (secondarily lost in some taxa) (Miller 2001). Miller (2001) classified Hydronebrius in the Agabinae, despite the latter's lack of the aforementioned setal series, due to its similar genitalia to derived members of the Agabini; its relationship to the Agabini requires investigation. The genus Hydrotrupes has also been excluded from Agabinae on the basis of the presence of open mandibular grooves in its larva (in contrast to the closed grooves found in other Agabinae and most other Dytiscidae except Copelatinae) but this separation was not supported by Miller (2001). The type genus Agabus is a primarily Holarctic group of species with a continuous bead along the anterior clypeal margin.

<==Agabinae
    |--Hydronebrius [Hydronebriini] M01
    `--Agabini M01
         |  i. s.: Andonectes M01
         |--Hydrotrupes palpalis M01
         `--+--+--Agametrus M01
            |  `--Leuronectes M01
            |       |--L. curtulus M01
            |       `--L. gaudichaudii M01
            |--+--Ilybius M01
            |  `--+--‘Agabus’ ambiguus M01
            |     |--‘Agabus’ bifarius M01
            |     |--‘Agabus’ disintegratus M01
            |     `--‘Agabus’ punctulatus M01
            |--Platynectes Regimbart 1878 M01, LM87 (see below for synonymy)
            |    |--*P. decemnotatus (Aubé 1838) [=Agabus decemnotatus] LM87
            |    |--P. aenescens Sharp 1882 [incl. P. subaenescens Lea 1899] LM87
            |    |--P. australicus Guéorguiev 1972 LM87
            |    |--P. bakewelli (Clark 1863) [=Agabus bakewelli; incl. P. obscurus Sharp 1882] LM87
            |    |--P. beroni S86
            |    |--P. brownei Guéorguiev 1972 [=P. (*Australonectes) brownei] LM87
            |    |--P. chapmani S86
            |    |--P. darlingtoni Guéorguiev 1972 [incl. P. multimaculatus Watts 1978] LM87
            |    |--P. decempunctatus (Fabricius 1775) (see below for synonymy) LM87
            |    |--P. laurianus Watts 1978 LM87
            |    |--P. magellanicus (Babington 1841) [=Colymbetes magellanicus, P. (*Neoplatynectes) magellanicus] LM87
            |    |--P. monostigma (Hope 1842) [=Colymbetes monostigma; incl. Agabus latissimus Clark 1863] LM87
            |    |--P. octodecimmaculatus (Macleay 1825) [=Colymbetes octodecimmaculatus] LM87
            |    |--P. reticulosus (Clark 1863) [=Agabus reticulosus; incl. P. limbatus Sharp 1882] LM87
            |    `--P. tasmaniae (Clark 1863) (see below for synonymy) LM87
            `--+--+--Agabinus glabrellus M01
               |  `--Platambus M01
               |       |--P. (Platambus) maculatus M01
               |       `--P. (Anagabus) lineatus M01
               `--Agabus Leach 1817 M01
                    |  i. s.: A. heydeni RB02
                    |         A. paludosus C01
                    |         A. ragazzii G89
                    |--A. spinipes M01
                    `--+--+--+--A. cordatus M01
                       |  |  `--A. seriatus M01
                       |  `--+--A. punctatus M01
                       |     |--A. semivittatus M01
                       |     `--A. tristis M01
                       `--+--A. unguicularis M01
                          |--+--A. antennatus M01
                          |  `--A. crassipes [=Carrhydrus crassipes] M01
                          |--+--A. approximatus M01
                          |  |--A. griseipennis M01
                          |  `--A. obliteratus M01
                          `--+--A. japonicus M01
                             `--+--A. minnesotensis M01
                                `--A. regularis M01

Platynectes Regimbart 1878 M01, LM87 [incl. Australonectes Guéorguiev 1972 LM87, Neoplatynectes Guéorguiev 1972 LM87]

Platynectes decempunctatus (Fabricius 1775) [=Dytiscus decempunctatus; incl. Agabus lugubris Blanchard 1853, A. mastersii Macleay 1871, Lancetes ocularis Lea 1895, Platynectes polygrammus Régimbart 1899, Agabus spilopterus Germar 1848] LM87

Platynectes tasmaniae (Clark 1863) [=Agabus tasmaniae; incl. P. daemeli Sharp 1882, P. gagatinus Lea 1899] LM87

*Type species of generic name indicated

REFERENCES

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45–92.

[RB02] Ribera, I., R. G. Beutel, M. Balke & A. P. Vogler. 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B—Biological Sciences 269: 2351–2356.

[S86] Spangler, P. J. 1986. Insecta: Coleoptera. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 622–631. E. J. Brill/Dr W. Backhuys: Leiden.

Last updated: 5 February 2018.