Male Oodes helopioides, copyright U. Schmidt.

Belongs within: Harpalinae.
Contains: Panagaeitae, Chlaenius.

The Oodini are a group of ground beetles characterised by a twisted fold near the apex of the elytral epipleura (Bell 1990).

Characters (from Bell 1990): Mandibles usually pointed, not strongly curved; scrobal seta absent. Eye usually large; one orbital seta over each eye. Pronotum more or less flattened, broadest at or near base, basal impressions scarcely evident. Elytra glabrous; stria 8 much deeper than stria 7; interval 9 narrow, careniform; epipleura interrupted by twisted fold near apex. Metepimeron visible as scale-like structure overlapping abdominal sternum I at side.

<==Oodini [Oodinae] B90
    |  i. s.: Evolenes exarata B90
    |         Dercylinus impressus B90
    |         Stenomorphus B90
    |         Systolocranius validus B55
    |         Paroodes similatus B55
    |         Lonchosternus B55
    |           |--L. deceptor B55
    |           `--L. politus B55
    |--+--+--Anatrichis LeConte 1853 OM08, LM87
    |  |  |    |--*A. minutus (Dejean 1831) [=Oodes minutus] LM87
    |  |  |    |--A. australasiae Chaudoir 1882 [incl. Oodes pygmaeus Macleay 1888] LM87
    |  |  |    |--A. lilliputana (Macleay 1888) [=Oodes lilliputanus] LM87
    |  |  |    |--A. pusilla Sloane 1910 LM87
    |  |  |    `--A. sexstriata Sloane 1910 LM87
    |  |  `--Dercylus [Dercylini] OM08
    |  `--+--Adelopomorpha glabra OM08
    |     `--Panagaeitae OM08
    `--+--Stenocrepis elegans OM08
       `--+--Oodes Bonelli 1810 OM08, LM87
          |    |--*O. helopioides (Fabricius 1792) [=Carabus helopioides] LM87
          |    |--O. abaxoides Lucas 1846 E12
          |    |--O. amaroides OM08
          |    |--O. bostockii Castelnau 1867 LM87
          |    |--O. denisonensis Castelnau 1867 LM87
          |    |--O. fitzroyensis Macleay 1888 LM87
          |    |--O. froggatti Macleay 1888 LM87
          |    |--O. impressus Chaudoir 1882 LM87
          |    |--O. inornatus Castelnau 1867 [incl. O. proximus Castelnau 1867] LM87
          |    |--O. latior Csiki 1931 [=O. latus Castelnau 1867 non LaFerté 1851] LM87
          |    |--O. mauritanicus Lucas 1846 E12
          |    |--O. modestus Castelnau 1867 LM87
          |    |--O. oblongus Castelnau 1867 (see below for synonymy) LM87
          |    |--O. paroensis Castelnau 1867 LM87
          |    |--O. parviceps Sloane 1896 LM87
          |    |--O. trisulcatus Castelnau 1867 LM87
          |    `--O. waterhousei Castelnau 1867 [incl. O. interioris Castelnau 1867] LM87
          `--Chlaeniini [Chlaeniides, Chlaeniinae] OM08
               |--Chlaenius OM08
               |--Rhysotrachelus quadrimaculatus B55
               |--Ocybatus deyrollei B55
               |--Stenodinodes nepos B55
               |--Pachydinodes conformis B55
               |--Chlaenites capicola B55
               |--Pseudochlaeniellus paenulatus B55
               |--Callistoides B55
               |    |--C. hessei Basilewsky 1949 B55
               |    `--C. pulchellus B55
               |--Leptochlaenius varians B55
               |    |--L. v. varians B55
               |    |--L. v. angolanus Basilewsky 1955 B55
               |    `--L. v. verecundus B55
               |--Trachychlaenites B55
               |    |--T. gonioderus B55
               |    |    |--T. g. gonioderus B55
               |    |    `--T. g. nigrofemoralis B55
               |    `--T. obesus [incl. T. ovalipennis] B55
               |--Homalolachnus B55
               |    |--H. goossensi [incl. H. goossensi var. nigrocyanescens] B55
               |    `--H. lineatus B55
               |         |--H. l. lineatus B55
               |         `--H. l. tristis B55
               `--Chlaeniostenus B55
                    |--C. denticulatus B55
                    |    |--C. d. denticulatus B55
                    |    `--C. d. elatus B55
                    |--C. euryscopus B55
                    `--C. sulcipennis B55
                         |--C. s. sulcipennis B55
                         `--C. s. sulcatulus B55

Oodes oblongus Castelnau 1867 [incl. Coptocarpus planipennis Macleay 1878, C. riverinae Macleay 1873] LM87

*Type species of generic name indicated


[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

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