Hydrocanthus iricolor, photographed by Tom Murray.

Belongs within: Adephaga.
Contains: Dytiscidae.

The Dytiscoidea is a clade of predatory beetles united by the possession of extensively fused mesal metacoxal walls (Ribera et al. 2002). Though all dytiscoids are found in damp habitats, advanced swimming abilities through simultaneous strokes of the hind legs are only found in species of Noteridae and Dytiscidae. Hygrobiidae swim with alternate movements of the hind legs, while Amphizoa and Aspidytes do not swim.

    |  i. s.: Necronectulus S02
    |         Colymbothetis [Colymbothetidae] S02
    |         Parahygrobia [Parahygrobiidae] P02
    |           `--P. natans P02
    |         Liadytidae RB02
    |           |--Liadytes longus RB02, P02
    |           `--Memptus S02
    |--+--Amphizoa [Amphizoidae] RB02
    |  |    |--A. insolens RB02
    |  |    `--A. leconti RB02
    |  `--+--Aspidytes Ribera, Beutel et al. 2002 [Aspidytidae] RB02
    |     |    `--*A. niobe Ribera, Beutel et al. 2002 RB02
    |     `--+--Dytiscidae RB02
    |        `--Hygrobia [Hygrobiidae] RB02
    |             |  i. s.: H. tarda M01
    |             |--H. hermanni RB02
    |             `--+--H. australasiae RB02
    |                `--H. maculata RB02
    `--Noteridae RB02
         |  i. s.: Phreatodytes Ueno 1957 S86
         |           `--P. relictus Ueno 1957 S86
         |--Notomicrus tenellus RB02
         `--+--Noterus clavicornis RB02
            |--+--Mesonoterus crassicornis M01
            |  `--Pronoterus punctipennis M01, R89
            `--+--Suphis cimicoides RB02, M01
               `--+--Suphisellus RB02
                  |--Hydrocanthus M01
                  |    |--H. debilis R89
                  |    |--H. iricolor M01
                  |    |--H. laevigatus M01
                  |    `--H. paludimonstrus M01
                  `--Canthydrus M01
                       |--C. balzani Régimbart 1889 R89
                       |--C. crassicornis Régimbart 1889 R89
                       |--C. flavolineatus Régimbart 1889 R89
                       |--C. flavopictus Régimbart 1889 R89
                       |--C. grammicus R89
                       |--C. grammopterus Régimbart 1889 R89
                       |--C. grossus R89
                       |--C. infuscatus Régimbart 1889 R89
                       |--C. nigrinus M01
                       |--C. obscuripennis Régimbart 1889 R89
                       |--C. remator R89
                       |--C. rotundatus R89
                       `--C. sexnotatus Régimbart 1889 R89

*Type species of generic name indicated


[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45-92.

[P02] Ponomarenko, A. G. 2002. Superorder Scarabaeidea Laicharting, 1781. Order Coleoptera Linné, 1758. The beetles. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 164-176. Kluwer Academic Publishers: Dordrecht.

[R89] Régimbart, M. 1889. Enumération des Haliplidae, Dytiscidae et Gyrinidae recueillis par M.r le Prof. L. Balzan dans l'Amérique méridionale et description de quelques autres espèces voisines. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 256-268.

[RB02] Ribera, I., R. G. Beutel, M. Balke & A. P. Vogler. 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B – Biological Sciences 269: 2351-2356.

[S02] Sinitshenkova, N. D. 2002. Ecological history of the aquatic insects. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 388-426. Kluwer Academic Publishers: Dordrecht.

[S86] Spangler, P. J. 1986. Insecta: Coleoptera. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 622-631. E. J. Brill / Dr. W. Backhuys: Leiden.

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