Sunburst diving beetle Thermonectus marmoratus, from Insectarium de Montréal, René Limoges.

Belongs within: Dytiscinae.

The Aciliini are a clade of diving beetles, distinguished by the possession of bifid metatibial spurs, and extremely long gonocoxae with expanded and rounded apices and apicolateral margins concave (Miller 2003). Members of the Aciliini lay their eggs in cracks in wood, moss or other cryptic locations (Miller 2001).

    |  i. s.: Tikoloshanes eretiformis M01
    |         Aethionectes M01
    |           |--A. apicalis M01
    |           `--A. optatus M01
    |--Thermonectus M03
    |    |  i. s.: T. succinctus R89
    |    |--T. marmoratus M03
    |    `--+--T. basilaris M01
    |       |--T. circumscriptus M03
    |       |--T. margineguttatus M01
    |       |--T. nigrofasciatus M03
    |       |--T. nobilis M01
    |       `--T. ornaticollis M01
    `--+--Rhantaticus congestus M03
       `--+--Sandracottus bakewelli M03
          `--+--Graphoderus M03
             |    |  i. s.: G. occidentalis M01
             |    |         G. perplexus M01
             |    |--G. liberus M03
             |    `--+--G. fascicollis M03
             |       `--G. zonatus M03
             `--Acilius M03
                  |  i. s.: A. japonicus I92
                  |         A. kishii Nakane 1963 I92
                  |--A. sulcatus M03
                  `--A. (Acilius) M01
                       |  i. s.: A. (A.) abbreviatus M01
                       |--A. (A.) semisulcatus M03, M01
                       `--+--A. (A.) sylvanus M03, M01
                          `--A. (Homeodytes) mediatus M03, M01

*Type species of generic name indicated


[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45-92.

[M03] Miller, K. B. 2003. The phylogeny of diving beetles (Coleoptera: Dytiscidae) and the evolution of sexual conflict. Biological Journal of the Linnaean Society 79: 359-388.

[R89] Régimbart, M. 1889. Enumération des Haliplidae, Dytiscidae et Gyrinidae recueillis par M.r le Prof. L. Balzan dans l'Amérique méridionale et description de quelques autres espèces voisines. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 256-268.

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