Cossyphodini

Cossyphodes arnoldi, from Schawaller (2013).

Belongs within: Tenebrionidae.

The Cossyphodini are a group of specialised inquiline beetles found in association with ants. They are rarely collected but most diverse in Africa, with a smaller number of species known from southern Asia.

Characters (from Andreae 1961): Head more or less semicircular; deep antennal pouches present between cheek and throat plate. Antennae geniculate, 9- or 11-segmented, with 1- or 2-segmented club respective to segment number. Tarsal formula 5,5,4 or 5,4,4.

<==Cossyphodini [Cossyphodidae] B14
    |  i. s.: Esemephe tumi Steiner 1980 B14
    |--Cossyphodites Brauns 1901 [Cossyphoditinae] A61
    |    `--*C. woodroffei (Péringuey 1885) [=Cossyphodes woodroffei] A61
    |--Paramelloninae A61
    |    |--Paramellon sociale Waterhouse 1882 A61
    |    `--Cossyphodinus Wasmann 1899 A61
    |         `--C. indicus A61
    `--Cossyphodinae A61
         |--Paramellops Andreae 1961 A61
         |    |--*P. bewicki (Wollaston 1861) [=Cossyphodes bewickii; incl. C. beccarii Gestro 1872] A61
         |    `--P. voigti (Reichensperger 1925) [=Cossyphodes voigti] A61
         `--Cossyphodes Westwood 1850 A61
              |  i. s.: C. angolanus Basilewsky 1952 A61
              |         C. uviranus Basilewsky 1950 A61
              |--C. (Cossyphodes) A61
              |    |--*C. wollastoni Westwood 1850 A61
              |    `--C. braunsi Andreae 1961 A61
              |--C. (Allocossyphodes Andreae 1961) A61
              |    `--C. (*A.) arnoldi Brauns 1925 A61
              |--C. (Hypercossyphodes Andreae 1961) A61
              |    |--C. (*H.) vandami Andreae 1961 A61
              |    |--C. (H.) brincki Andreae 1961 A61
              |    |--C. (H.) godfreyi Andreae 1961 A61
              |    `--C. (H.) natalensis Andreae 1961 A61
              |--C. (Metacossyphodes Andreae 1961) A61
              |    `--C. (*M.) kundelungensis Basilewsky 1950 A61
              |--C. (Mimocossyphus) A61
              |    |--C. (*M.) minor Pic 1923 A61
              |    |--C. (M.) freudei Basilewsky 1957 A61
              |    |--C. (M.) fulvus Andreae 1961 A61
              |    |--C. (M.) matabele Andreae 1961 A61
              |    |--C. (M.) raffrayi Gestro 1874 A61
              |    |--C. (M.) reichenspergeri Andreae 1961 A61
              |    |--C. (M.) stevensoni Andreae 1961 A61
              |    `--C. (M.) wasmanni Reichensperger 1916 A61
              `--C. (Pachycossyphodes Andreae 1961) A61
                   |--C. (*P.) machadoi Basilewsky 1952 A61
                   `--C. (P.) pretoriensis Andreae 1961 A61

*Type species of generic name indicated

REFERENCES

[A61] Andreae, H. 1961. Coleoptera: Cossyphodidae. In: Hanström, B., P. Brinck & G. Rudebeck (eds) South African Animal Life: Results of the Lund University Expedition in 1950–1951 vol. 8 pp. 198–216. Almqvist & Wiksell: Uppsala.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).