Coffee bean weevil Araecerus fasciculatus, from PaDIL.

Belongs within: Coleoptera.

The Anthribidae are a group of weevils with a flattened rostrum retaining a visible labrum, and non-elbowed antennae, whose larvae feed on dead wood, galls and fungi (Britton 1970).

Characters (from Britton 1970): Rostrum, broad, short or elongate, flattened; antennae not elbowed, scape shorter than pedicel, short with three-segmented club, or elongate without club or with only slightly marked club; labrum visible; maxillae with lacinia present; maxillary palpi slender, four-segmented, flexible; mentum large, pedunculate, ligula large, sclerotised; gular sutures not visible; prothorax with lateral margins present on basal half; fore coxal cavities contiguous, coxae globular; mid coxal cavities broadly closed by sterna; abdomen with four sternites connate, apical sternite only movable; elytra with inner flange below shoulder; pygidium largely exposed; tarsi with third segment deeply bilobed, embraced by prominent angles of second; claws sometimes toothed beneath. Larva curved, fleshy, widest in middle; head usually exserted, hypognathous; antennae one-segmented or absent; legs absent or with reduced segments.

    |--Urodontinae [Urodontidae] B14
    |    |--Urodon pygmaeus C01
    |    |--Bruchela [Bruchelinae] B14
    |    |--Cercomorphus B14
    |    `--Urodontellus B14
    |         |--U. lilii (Fåhraeus 1839) B14
    |         |--U. vermiculatus B14
    |         `--U. vicinialilii B14
    |--Choraginae B14
    |    |--Choragini B14
    |    |    |--Apteroxenus globulosus Valentine 1979 B14
    |    |    `--Euxenulus B14
    |    `--Araecerus Schoenherr 1823 M94 [Araecerini B14]
    |         |--*A. coffeae (Fabricius 1801) [=Anthribus coffeae] M94
    |         |--A. fasciatus B49
    |         |--A. fasciculatus (de Geer 1775) B14 [incl. A. sambucinus MacLeay in Boisduval 1835 M87, B35]
    |         |--A. lutatus (Fairmaire 1849) [incl. A. vieillardi] M94
    |         `--A. palmaris (Pascoe 1882) [=Doticus palmaris] M94
    `--Anthribinae B14
         |  i. s.: Anthribus B14
         |           |--A. attelaboides [=Rhinomacer attelaboides] L02
         |           |--A. fasciatus B14
         |           |--A. latirostris L02
         |           |--A. nebulosus Forster 1770 B14
         |           |--A. rufipes [=Bruchus rufipes] L02
         |           |--A. scabrosus L02
         |           |--A. scapularis B14
         |           `--A. suturalis [=Bruchus suturalis] L02
         |         Toxonotus MW15
         |         Cacephatus Blackburn 1900 M94
         |           |--*C. sericeus Blackburn 1900 M94
         |           |--C. aucklandicus M94
         |           |--C. huttoni M94
         |           `--C. inornatus M94
         |         Euciodes Pascoe 1866 M94
         |           `--*E. suturalis Pascoe 1866 M94
         |         Eucorynus Schoenherr 1826 M94
         |           |--*E. crassicornis [=Anthribus crassicornis] M94
         |           `--E. mastersi Blackburn 1900 M94
         |         Phloeobius Schoenherr 1823 M94
         |           `--P. gigas (Fabricius 1775) M94 (see below for synonymy)
         |         Platystomos albinus GC-P14
         |--Exechesops [Zygaenodini] B14
         |    `--E. leucopis (Jordan 1928) B14
         `--Gymnognathini B14
              |--Xynotropis B14
              `--Lichenobius B14
                   |--L. littoralis Holloway 1970 B14
                   |--L. maritimus B14
                   `--L. silvicola B14

Anthribidae incertae sedis:
  Notochoragus crassus (Sharp 1876) E02
  Anthribites rechenbergi Kolbe 1888 (n. d.) Z02
  Brachytarsus pantherinus Lucas 1846 A71, E12
  Phymatus phymatodes WFS04
  Pleosporius bullatus WFS04
  Sharpius brouni [=Anthribus brouni] B88
  Euxenus L88
  Cyptoxenus L88
  Cacorhinus lateripictus LB91
  Ancyclotropis waterhousei LB91
  Xenocerus M87
    |--X. arciferus Blanch 1853 M86
    `--X. leucogrammus Mots. 1874 M87
  Litocerus M86
    |--L. fasciatus Macleay 1886 M86
    |--L. parvulus Macleay 1886 M86
    |--L. perplexus M86
    `--L. subconvexus Macleay 1886 M86
  Nessiara M86
    |--N. irrorata Macleay 1886 M86
    `--N. unituberculata Macleay 1886 M86
  Bythoprotus lineatus M87
  Ancylotropis waterhousei M87
  Genethila retusa M87
  Macrotrichius niveinasus Mots. 1874 M87
  Ethneca bakewelli M87
  Tropidoderes M87
    |--T. albuginosus Erichs. 1842 M87
    `--T. musivus Erichs. 1842 M87
  Ecelonerus M87
    |--E. albopictus Pascoe 1860 M87
    |--E. insularis M87
    `--E. subfasciatus M87
  Baseotropis M87
    |--B. ingrata Pascoe 1859 M87
    |--B. peregrina Pascoe 1859 M87
    `--B. solitaria Pascoe 1860 M87
  Ozotomerus Z93
    |--O. bipunctatus Z93
    `--O. waterhousei (Pascoe 1859) [=Dipieza waterhousei] M87
  Cratoparis M87
    |--C. callosus M87
    `--C. ceroderes M87
  Teratanthribus monstrosus Z93
  Basitropis Z94
  Autotropis modesta M94
  Dendrotrogus colligens B70
  Doticus pestilans B70

Phloeobius gigas (Fabricius 1775) M94 [=Anthribus gigas M94; incl. A. griseus M94, *Phloeobius griseus M94, M87]

*Type species of generic name indicated


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[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

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[E02] Emberson, R. M. 2002. The beetle (Coleoptera) fauna of the Chatham Islands: additions and corrections. New Zealand Entomologist 25: 69–77.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GC-P14] Gillett, C. P. D. T., A. Crampton-Platt, M. J. T. N. Timmermans, B. H. Jordal, B. C. Emerson & A. P. Vogler. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution 31 (8): 2223–2237.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385–409.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera" (continued). Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 183–204.

[M87] Masters, G. 1887. Catalogue of the described Coleoptera of Australia. Part VI. Proceedings of the Linnean Society of New South Wales, series 2, 1 (4): 979–1036.

[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[WFS04] Winks, C. J., S. V. Fowler & L. A. Smith. 2004. Invertebrate fauna of boneseed, Chrysanthemoides monilifera ssp. monilifera (L.) T. Norl. (Asteraceae: Calenduleae), an invasive weed in New Zealand. New Zealand Entomologist 27: 61–72.

[Z02] Zherikhin, V. V. 2002. Pattern of insect burial and conservation. In: Rasnitsyn, A. P.,  & D. L. J. Quicke (eds) History of Insects pp. 17–63. Kluwer Academic Publishers: Dordrecht.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.

[Z94] Zimmerman, E. C. 1994. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May. CSIRO Australia.

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