Cossoninae

Cossonus linearis, copyright Siga.

Belongs within: Curculionidae.

The Cossoninae are a group of elongate-bodied weevils that are borers in dead or dying wood (Marvaldi & Lanteri 2005).

Characters (from Marvaldi & Lanteri 2005): Body elongate, parallel-sided, depressed. Rostrum often shorter than prothorax. All legs usually with uncinate tibiae; front tibiae with grooming device consisting of long comb of setae on apical third to half; hind tibiae lacking apical comb of setae. Tarsite 3 as wide as tarsite 2.

<==Cossoninae [Cossonides] GC-P14
    |--Acamptini B14
    |    |--Acamptella B14
    |    |--Trachodisca B14
    |    |--Pseudacamptopsis B14
    |    |--Acamptopsis B14
    |    |--Choerorrhynchus B14
    |    |--Menares B14
    |    |--Prionarthrus B14
    |    `--Acamptus [incl. Glyphostethus] Z93
    |         |--A. cancellatus B14
    |         `--A. rigidus LeConte 1876 B14
    |--Cossonini M94
    |    |--Mesites Schoenherr 1838 M94
    |    |    |--*M. pallidipennis Boheman 1837 M94, HF03
    |    |    `--M. cunipes Boheman 1837 HF03
    |    `--Cossonus D09
    |         |--C. albertisi Pascoe 1885 Mas86
    |         |--C. basalis Pascoe 1885 Mac86
    |         |--C. excavatus Pascoe 1885 Mas86
    |         |--C. fossatus ML05
    |         |--C. impressifrons Boheman 1838 D09
    |         |--C. indigens Pascoe 1885 Mas86
    |         |--C. linearis L02
    |         |--C. praeustus Mas86
    |         `--C. simsoni LB91
    |--Pentarthrum Wollaston 1854 [Pentarthrini] M94
    |    |--*P. huttoni Wollaston 1854 M94
    |    |--P. elumbe GC-P14
    |    |--P. helmsianum H15
    |    |--P. nigrum Woll. 1873 Mas86
    |    `--P. zealandicum Wollaston 1873 M94
    |--Araucariini M94
    |    |--Araucarius GE05
    |    `--Xenocnema Wollaston 1873 M94
    |         `--*X. spinipes Wollaston 1873 M94
    `--Rhyncolini M94
         |--Rhyncolus knowltoni W86
         `--Phloeophagosoma Wollaston 1873 M94
              |--*P. minutum Wollaston 1873 M94
              |--P. aesculi M94
              |--P. tenuis M94
              `--P. thoracicum Wollaston 1874 M94

Cossoninae incertae sedis:
  Choerorhinus squalidus Fairmaire 1857 HF03
  Pselactus spadix (Herbst 1795) HF03
  Pseudomimus avocadi Folwaczny 1973 HF03
  Phoenomerus exilis Pascoe 1872 Mas86
  Notiomimetes pascoei Woll. 1873 Mas86
  Cossonideus Mas86
    |--C. lineola (Fabricius 1775) [=Curculio lineola] Z93
    `--C. pascoei Woll. 1873 Mas86
  Halorhynchus caecus Woll. 1873 Mas86
  Thaumastophasis oculatus Woll. 1873 Mas86
  Stereomimetes crassicornis Woll. 1873 Mas86
  Notiosomus Mas86
    |--N. australis Woll. 1873 Mas86
    |--N. congener Woll. 1873 Mas86
    `--N. major Woll. 1873 Mas86
  Homalotrogus arctatus Pascoe 1885 Mas86
  Aphanocorynes depressus Woll. 1873 Mas86
  Pentamimus Mas86
    |--P. canaliculatus Woll. 1873 Mas86
    |--P. rhyncholiformis Woll. 1873 Mas86
    `--P. suffusus Woll. 1873 Mas86
  Isotrogus bilineatus Pascoe 1885 Mas86
  Hyponotus subpubescens Woll. 1873 Mas86
  Orthotemnus disparilis Pascoe 1885 Mas86
  Stereoborus inductus Pascoe 1885 Mas86
  Dryotribus Z93
  Aphyoda Z93
  Himatium [incl. Caulosomus] Z93
  Eiratus M94
  Exeiratus M94

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[D09] Davis, S. R. 2009. Morphology of Baridinae and related groups (Coleoptera, Curculionidae). ZooKeys 10: 1–136.

[GC-P14] Gillett, C. P. D. T., A. Crampton-Platt, M. J. T. N. Timmermans, B. H. Jordal, B. C. Emerson & A. P. Vogler. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution 31 (8): 2223–2237.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HF03] Halperin, J., & J. Fremuth. 2003. Contribution to the knowledge of Curculionoidea (Coleoptera) and their host plants in Israel. Zoology in the Middle East 29: 93–100.

[H15] Hedley, C. 1915. Presidential address. Journal and Proceedings of the Royal Society of New South Wales 49 (1): 1–77, pls 1–7.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[Mac86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera" (continued). Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 183–204.

[ML05] Marvaldi, A. E., & A. A. Lanteri. 2005. Key to higher taxa of South American weevils based on adult characters (Coleoptera, Curculionoidea). Revista Chilena de Historia Natural 78: 65–87.

[Mas86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part V. Proceedings of the Linnean Society of New South Wales, series 2, 1 (3): 585–686.

[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.

[W86] Wood, S. L. 1986. A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs 10: 1–126.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.