Pristoderus antarcticus, copyright Udo Schmidt.

Belongs within: Tenebrionoidea.

The Zopheridae, ironclad beetles, are a group of beetles usually found living under bark of dead or dying trees.

Characters (from Ironclad ID): Antenna 9–11 segmented with a usually abrupt, 1–3 segmented club; antennal insertions concealed from above; mesocoxal cavities closed; trochanters heteromeroid; tarsal formula 4-4-4 or 5-5-4; male genitalia with tenebrionoid aedeagus.

Zopheridae B14
    |--Zopherinae B14
    |    |--Usechini B14
    |    |    |--Usechus B14
    |    |    `--Usechimorpha B14
    |    |         |--U. barberi B14
    |    |         `--U. montanus Doyen & Lawrence 1979 B14
    |    `--Zopherini B14
    |         |--Zopherus chilensis Gray 1823 B14
    |         |--Zopher B14
    |         |--Zopherosis georgii White 1859 B14, M86
    |         `--Scoriaderma B14
    `--Colydiinae [Colydiidae] B14
         |  i. s.: Enarsus WH02
         |         Ditoma crenata C01
         |         Rhopalocerus LB91
         |         Sparactus elongatus LB91
         |         Pseudendestes LB91
         |           |--P. australis LB91
         |           `--P. robertsi LB91
         |         Todinia LB91
         |         Ablabus LB91
         |         Dryptops LB91
         |         Nematidium [Nematidiini] B14
         |           `--N. filiforme LeConte 1863 B14
         |--Phellopsis MW15
         `--+--Pycnomerus helmsi MW15, H15
            `--Synchitini B14
                 |--Syncalus B14
                 |--Isotarphius B14
                 `--Pristoderus MW15
                      |--P. antarcticus (White 1846) B14
                      |--P. plagiatus (Broun 1911) E02
                      `--P. saccharatus (Pascoe 1870) [=Byrsax saccharatus] MB08

Zopheridae incertae sedis:
  Notocoxelus E02
  Heterargus ED04
  Cotulades LB91
    |--C. fascicularis Pascoe 1860 M86
    |--C. funerosa Hope 1845 M86
    `--C. leucospila Hope 1845 M86
  Latometus pubescens Erichs. 1842 LB91, M86
  Docalis LB91
    |--D. degener Pascoe 1860 M86
    `--D. exoletus Pascoe 1860 M86

*Type species of generic name indicated


[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E02] Emberson, R. M. 2002. The beetle (Coleoptera) fauna of the Chatham Islands: Additions and corrections. New Zealand Entomologist 25: 69–77.

[ED04] Ewers, R. M., & R. K. Didham. 2004. A first assessment of New Zealand beetles restricted to large forest areas. Weta 27: 33–40.

[H15] Hedley, C. 1915. Presidential address. Journal and Proceedings of the Royal Society of New South Wales 49 (1): 1–77, pls 1–7.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[MB08] Matthews, E. G., & P. Bouchard. 2008. Tenebrionid Beetles of Australia: Descriptions of tribes, keys to genera, catalogue of species. Australian Biological Resources Study: Canberra.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[WH02] Worthy, T. H., & R. N. Holdaway. 2002. The Lost World of the Moa: Prehistoric life of New Zealand. Indiana University Press: Bloomington (Indiana).

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