Nemonychidae

Dorsal and lateral views of Notomacer araucariae, from Legalov (2017). Scale bar = 1 mm.

Belongs within: Coleoptera.

The Nemonychidae, pine flower weevils, are a group of weevils with larvae feeding on pollen of conifers. Adults are distinguished from other weevils by the combination of a free labrum, visible at the level of the clypeus, together with a pronotum lacking lateral carinae, narrowly separated mid coxae and all free abdominal ventrites.

<==Nemonychidae [Eobelidae, Rhinomaceridae, Rhinomacerides] B14
    |  i. s.: Brenthorrhinus mirabilis P02
    |         Pagomacer M94
    |         Nannomacer M94
    |         Rhynchitomacer M94
    |         Eobelus longipes RJ93
    |--Nemonyx [Nemonychinae] MW15
    |    `--N. lepturoides M94
    |--Cimberidinae GC-P14
    |    |--Doydirhynchus austriacus GC-P14
    |    `--Cimberis B14 [incl. Rhinomacer RD77]
    |         |--C. attelaboides B14
    |         |--C. australiae [=Rhinomacer australiae] B70
    |         |--‘Rhinomacer’ curculionoides L02
    |         |--C. decipiens B14
    |         |--C. elongata (LeConte 1876) B14
    |         |--C. pallipennis B14
    |         `--C. pilosa M94
    `--Rhynorhynchinae MW15
         |  i. s.: Rhynchitomacerinus MW15
         |--Rhinorhynchini Z94
         |    |--Auletanus Z94
         |    |--Auletulus Z94
         |    |--Salacus Z94
         |    |--*Basiliogeus prasinus M94
         |    |--*Basiliorhinus araucariae M94
         |    `--Rhinorhynchus [=Listrorhinus] Z94
         |         `--*R. rufulus [=Rhinomacer rufulus, *Listrorhinus rufulus; incl. Rhinorhynchus zealandicus] Z94
         `--Mecomacerini B14
              |--Brarus mystes Kuschel 1997 B14
              |--Mecomacer B14
              |--Araucomacer B14
              |--*Aragomacer leai M94
              |--*Eutactobius puellus M94
              |--Bunyaeus M94
              |    |--*B. monteithi M94
              |    `--B. eutactae M94
              `--Notomacer M94
                   |--*N. araucariae M94
                   |--N. australiae M94
                   |--N. barbatulus M94
                   |--N. caledonicus M94
                   |--N. reginae M94
                   |--N. uniformis M94
                   `--N. zimmermani M94

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[GC-P14] Gillett, C. P. D. T., A. Crampton-Platt, M. J. T. N. Timmermans, B. H. Jordal, B. C. Emerson & A. P. Vogler. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution 31 (8): 2223–2237.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[P02] Ponomarenko, A. G. 2002. Superorder Scarabaeidea Laicharting, 1781. Order Coleoptera Linné, 1758. The beetles. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 164–176. Kluwer Academic Publishers: Dordrecht.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10^th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[Z94] Zimmerman, E. C. 1994. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May. CSIRO Australia.

Last updated: 15 November 2020.