Bembidion quadrimaculatum, copyright Udo Schmidt.

Belongs within: Bembidiinae.

The Bembidiina are a group of small carabids with reduced terminal palpomeres. It includes the large genus Bembidion with over 1200 species of relatively small ground beetles, most diverse in northern temperate regions.

<==Bembidiina R-JC01
    |--Notaphus Stephens 1828 R-JC01, LM87
    |    |--N. varius (Olivier 1795) [=Carabus varius, Bembidion (*Notaphus) varium] LM87
    |    |--N. brullei (Gemminger & Harold 1868) (see below for synonymy) LM87
    |    `--N. convergens [=Austronotaphus convergens] R-JC01
    |--Peryphus Stephens 1828 R-JC01, LM87
    |    |--*P. litoralis (Olivier 1795) [=Carabus litoralis, Bembidion (Peryphus) litorale] LM87
    |    |--P. bandotaro (Morita 1991) [=Bembidion (Peryphus) bandotaro] L98
    |    |--P. chilensis [=Chilioperyphus chilensis] R-JC01
    |    |--P. postae (Csiki 1901) [=Bembidium (Peryphus) postae] C01
    |    `--P. tetracolus (Say 1823) [=Bembidium tetracolum, Bembidion (Peryphus) tetracolum] LM87
    `--Bembidion Latreille 1802 LM87
         |  i. s.: B. adustum C01
         |         B. aegyptiacum G89
         |         B. algiricum Lucas 1846 E12
         |         B. andreae C01
         |         B. articulatum C01
         |         B. assimile C01
         |         B. dentellum (Thunberg 1787) VPB11
         |         B. dives Lucas 1846 E12
         |         B. foveum C01
         |         B. gibberosum Lucas 1846 E12
         |         B. infuscatum C01
         |         B. lampros (Herbst 1784) VPB11 [incl. B. lampros var. properans C01]
         |         B. levettei OM08
         |           |--B. l. levettei OM08
         |           `--B. l. carrianum OM08
         |         B. mauritanicum Lucas 1846 E12
         |         B. numidicum Lucas 1846 E12
         |         B. octomaculatum C01
         |         B. perspicuum HR11
         |         B. pulchellum Lucas 1846 E12
         |         B. quadriguttatum [=Carabus quadriguttatus] L02
         |         B. quadripustulatum C01
         |         B. semilunium L98
         |         B. striatum C01
         |         B. vicinum Lucas 1846 E12
         |--*B. (Bembidion) quadrimaculatum (Linnaeus 1761) [=Cicindela quadrimaculata] LM87
         |--B. (Ananotaphus Nekolitzky 1931) LM87
         |    |--B. (*A.) errans Blackburn 1888 [=Bembidium errans] LM87
         |    |--B. (A.) blackburni Csiki 1928 [=Bembidium dubium Blackburn 1888 non Heer 1841] LM87
         |    `--B. (A.) proprium Blackburn 1888 [=Bembidium proprium] LM87
         |--B. (Desarmatocillenus Netolitzky 1942) LM87
         |    |--B. (*D.) yokohamae (Bates 1883) [=Cillenus yokohamae] LM87
         |    |--B. (D.) adelaidae Lindroth 1980 LM87
         |    |--B. (D.) albovirens (Sloane 1903) [=Cillenus albovirens] LM87
         |    `--B. (D.) mastersi Sloane 1895 (see below for synonymy) LM87
         |--B. (Notaphocampa Netolitzky 1914) [incl. Notaphomimus Netolitzky 1931] LM87
         |    |--B. (*N.) niloticum Dejean 1831 LM87
         |    `--B. opulentum Nietner 1858 [=Bembidium opulentum, Bembidion (*Notaphocampa) opulentum] LM87
         |         |--B. o. opulentum LM87
         |         `--B. o. riverinae Sloane 1895 [=Bembidium riverinae] LM87
         |--B. (Pekinium Csiki 1901) C01
         |    `--B. (*P.) chinense Csiki 1901 C01
         `--B. (Philochthus Stephens 1828) [incl. Sloanephila Netolitzky 1931] LM87
              |--B. (*P.) biguttatum (Fabrcius 1779) LM87 (see below for synonymy)
              `--B. (P.) jacksoniense Guérin-Méneville 1830 (see below for synonymy) LM87

Bembidion (Desarmatocillenus) mastersi Sloane 1895 [=Bembidium mastersi; incl. Bembidium flavescens Macleay 1864 (n. n.)] LM87

Bembidion (*Philochthus) biguttatum (Fabrcius 1779) LM87 [=Carabus biguttatus LM87; incl. B. biguttatum var. inoptatum G89]

Bembidion (Philochthus) jacksoniense Guérin-Méneville 1830 [=Bembidion (Leia) jacksoniensis, Bembidion (*Sloanephila) jacksoniense; incl. Bembidium ocellatum Blackburn 1888, Bembidium subviride Macleay 1871] LM87

Notaphus brullei (Gemminger & Harold 1868) [=Bembicidium brullei, Bembidion (Notaphus) brullei, Bembidion variegatum Brullé 1838 non Say 1823] LM87

*Type species of generic name indicated


[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.

[L98] Lafer, G. S. 1998. Supplementary accounts of the ground-beetles fauna (Coleoptera, Carabidae) of the Southern Kuril Islands. Far Eastern Entomologist 59: 19–20.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

Last updated: 21 February 2018.

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