Meloidae

Female Cerocoma schaefferi, copyright Siga.


Belongs within: Tenebrionoidea.
Contains: Mylabris, Zonitis, Meloe.

The Meloidae, blister beetles, are a group of soft-bodied beetles whose larvae are often parasitoids on bees.

Characters (from Lawrence & Britton 1991): Elongate, moderately convex, soft-bodied beetles, often bicoloured with red or yellow and black or blue, sometimes uniformly red or metallic blue or violet. Body subglabrous or clothed with short, decumbent hairs; head strongly constricted behind eyes to form narrow neck; antenna 11-segmented, filiform or serrate. Prothorax usually narrowed anteriorly, lacking lateral carinae: fore coxae projecting, trochantin exposed; fore coxal cavities open internally and externally. Tarsal formula 5-5-4, tarsal claws pectinate with blade-like process beneath each claw. Abdomen with six ventrites. Larvae of three types: minute, fusiform, well-sclerotised triungulin with long legs; grub-like, ectoparasitic larva, lightly sclerotised, with short legs and hypognathous head: coarctate form with reduced mouth-parts and appendages.

Meloidae [Cantharidiae]
    |--Horia [Horiales, Horiinae] LB91
    |    |--H. maculata L02
    |    |--H. mira LB91
    |    `--H. testacea L02
    |--Nemognathinae B14
    |    |--Nemognatha chrysomelina (Fabricius 1775) (see below for synonymy) G-PR05
    |    `--Stenodera [Stenoderini] B14
    |         |  i. s.: S. caucasica B14
    |         `--S. (Stenoderina) B14
    |              |--S. (S.) palaestina B14
    |              `--S. (S.) puncticollis (Chevrolat 1834) B14
    |--+--Zonitis LB91
    |  `--Palaestra LB91
    |       |--P. eucera Fairm. 1877 M86
    |       |--P. platycera Fairm. 1880 M86
    |       |--P. quadrifoveata Fairm. 1880 M86
    |       |--P. rubripennis M86
    |       `--P. rufocincta Fairm. 1880 M86
    `--Meloinae B14
         |--Meloe G-PR05
         |--Lytta [Lyttini] B14
         |    |--L. magister C96
         |    `--L. vesicatoria (Linnaeus 1758) (see below for synonymy) G-PR05
         |         |--L. v. vesicatoria B14
         |         |--L. v. freudei B14
         |         |--L. v. heydeni B14
         |         |--L. v. moreana B14
         |         `--L. v. togata B14
         |--Epicauta [Epicautini] B14
         |    |--E. atrata (Fabricius 1775) B14
         |    |--E. lemniscata BB68
         |    |--E. megalocephala [incl. E. megalocephala var. maura] C01b
         |    |--E. pardalis B14
         |    |--E. pennsylvanica B14
         |    |--E. rufidorsum (Goeze 1777) (see below for synonymy) G-PR05
         |    |--E. (Macrobasis) segmenta (Say 1824) [incl. Macrobasis cinctothorax Dugès 1889] G-PP-OS05
         |    `--E. sibirica C01a
         `--Cerocoma B14
              |  i. s.: C. muehlfeldi Gyllenhal 1817 G-PR05
              |         C. schreberi Fabricius 1781 G-PR05
              |         C. vahli Fabricius 1787 G-PR05
              `--C. (Cerocoma) B14
                   |--C. (C.) bernhaueri B14
                   |--C. (C.) dahli B14
                   |--C. (C.) prochaskana B14
                   |--C. (C.) schaefferi (Linnaeus 1758) B14 (see below for synonymy)
                   `--C. (C.) simplicicornis B14

Meloidae incertae sedis:
  Hornia A71
    |--H. gigantea R35
    `--H. minutipennis A71
  Cyaneolytta Péringuey 1909 G-PR05
    `--C. granulipennis (Laporte de Castelnau 1840) [=Lytta granulipennis] G-PR05
  Cabalia segetum (Fabricius 1792) [=Cantharis segetum, Lytta segetum] G-PR05
  Muzimes collaris (Fabricius 1787) [=Alosimus collaris, Halosimus kollaris (l. c.)] G-PR05
  Croscherichia G-PR05
    |--C. paykulli (Billberg 1813) (see below for synonymy) G-PR05
    `--C. sanguinolenta B14
  Physomeloe corallifer (Germar 1818) [=Meloe corallifer; incl. M. corallifer var. evae Flach 1907] G-PR05
  Leptopalpus rostratus (Fabricius 1792) [=Nemognatha rostrata, Zonitis rostratus] G-PR05
  Cysteodemus wislizeni C90
  Coryna Billberg 1813 G-PR05
    `--C. ocellata (Oliv. 1790) [=Cerocoma ocellata] G89
  Apalus G-PR05
    |--A. bimaculatus (Linnaeus 1761) G-PR05 [=Zonitis bimaculata L02]
    `--A. guerini (Mulsant 1858) [=Hapalus guerini] G-PR05
  Lagorina G-PR05
    |--L. scutellata (Laporte de Castelnau 1840) [=Cantharis scutellata] G-PR05
    `--L. sericea (Waltl 1835) (see below for synonymy) G-PR05
  Actenodia G-PR05
    |--A. billbergi (Gyllenhal 1817) (see below for synonymy) G-PR05
    `--A. distincta (Chevrolat 1837) [incl. Coryna distincta var. andalusiaca Pic 1916] G-PR05
  Berberomeloe G-PR05
    |--B. insignis (Charpentier 1818) [=Meloe majalis var. insignis] G-PR05
    `--B. majalis (Linnaeus 1758) G-PR05 (see below for synonymy)
  Stenoria G-PR05
    |--S. analis Schaum 1859 [=Apalus analis; incl. Hapalus analis var. acutipennis Fairmaire 1881] G-PR05
    `--S. apicalis (Latreille 1802) [=Hapalus apicalis, Sitaris apicalis] G-PR05
  Zonabris Harold 1879 G-PR05
    |--Z. maculata C01a
    |--Z. quatuordecimpunctata C01b
    `--Z. splendidula C01a
  Sitaris Latreille 1802 L02
    |--S. muralis (Foerster 1771) G-PR05 (see below for synonymy)
    |--S. rufipennis Küster 1849 G-PR05
    `--S. solieri Pecchioli 1839 G-PR05
  Oenas Latreille 1802 L02
    |--*O. atratus [=Lytta atrata] L02
    |--O. afer (Linnaeus 1758) G-PR05
    |--O. crassicornis (Illiger 1800) G-PR05
    |--O. fusicornis Abeille de Perrin 1880 [incl. O. hispanus Abeille de Perrin 1880] G-PR05
    `--O. ruficollis Olivier 1811 G-PR05
  Euzonitis G-PR05
    |--E. haroldi (Harold 1870) [=E. bipunctata var. haroldi, Zonitis quadripunctata var. haroldi] G-PR05
    |--E. paulinae (Mulsant & Rey 1858) [=Zonitis paolinae (l. c.)] G-PR05
    |--E. quadrimaculata (Pallas 1782) (see below for synonymy) G-PR05
    |--E. sexmaculata (Olivier 1789) (see below for synonymy) G-PR05
    `--E. terminata (Abeille de Perrin 1880) G-PR05
  Hycleus Latreille 1829 [incl. Gorrizia Pardo Alcaide 1954, Tigrabris Kuzin 1954] G-PR05
    |--H. dufourii (Graells 1849) [=Mylabris dufourii, Zonabris dufouri] G-PR05
    |--H. duodecimpunctatus (Olivier 1811) (see below for synonymy) G-PR05
    |--H. fuscus (Olivier 1811) [=Mylabris fusca] G-PR05
    |--H. polymorphus (Pallas 1771) (see below for synonymy) G-PR05
    |--H. praeustus (Fabricius 1792) [=Zonabris praeusta] G-PR05
    |--H. scutellatus (Rosenhauer 1856) (see below for synonymy) G-PR05
    `--H. wagneri (Chevrolat 1837) (see below for synonymy) G-PR05
  Mylabris G-PR05
  Iselma MW15
  Tmesidera C91
    |--T. assimilis Hope 1842 M86
    |--T. rubricollis Hope 1842 M86
    |--T. rufipennis Westw. 1841 M86
    `--T. violacea Hope 1842 M86

Nomen nudum: Oenas villiger Salvañá Comas 1870 G-PR05

Actenodia billbergi (Gyllenhal 1817) [=Coryna billbergi, Dices billbergi, Hycleus bilbergi, Mylabris bilbergi, Zonabris bilbergi] G-PR05

Berberomeloe majalis (Linnaeus 1758) G-PR05 [=Meloe majalis G-PR05; incl. M. fissicornis Górriz Muñoz 1882 G-PR05, M. laevigata Olivier 1792 G-PR05, M. majalis var. maculifrons Lucas 1847 G-PR05, E12, M. majalis var. sanguineus Escherich 1889 G-PR05]

Cerocoma (Cerocoma) schaefferi (Linnaeus 1758) B14 [=C. schafferi (l. c.) G-PR05; incl. C. wahlii var. chalybaeiventris Chevrolat 1838 G-PR05, C. schaefferi var. orensis Reitter 1913 G-PR05]

Croscherichia paykulli (Billberg 1813) [=Mylabris paykulii; incl. M. circumflexa Chevrolat 1837, Zonabris circumflexa] G-PR05

Epicauta rufidorsum (Goeze 1777) [incl. Cantharis dubia Olivier 1790, E. dubia, Lytta verticalis Illiger 1803, E. verticalis] G-PR05

Euzonitis quadrimaculata (Pallas 1782) [incl. Zonitis auricoma var. bombycina Escherich 1896, E. bipunctata ab. immaculata Escherich 1891, Z. quadripunctata var. korbi Escherich 1890, Z. bipunctata ab. korbi, Z. quadripunctata (Fabricius 1801), Z. quadripustulata (l. c.)] G-PR05

Euzonitis sexmaculata (Olivier 1789) [=Zonitis sexmaculata; incl. E. sexmaculata ab. bipunctata Ragusa 1882, E. sexmaculata ab. discolor Escherich 1891, Zonitis fenestrata Pallas 1781, E. sexmaculata ab. lunata Tauscher 1812, E. sexmaculata ab. pedemontana Pick 1901] G-PR05

Hycleus duodecimpunctatus (Olivier 1811) [=Mylabris duodecimpunctata, Zonabris duodecimpunctata; incl. M. cyanescens, Z. duodecimpunctata var. fasciata De la Fuente 1910, Z. duodecimpunctata var. massiliensis Pic 1902] G-PR05

Hycleus polymorphus (Pallas 1771) [incl. Mylabris cichorii (Schrank 1781), M. floralis (Pallas 1781), Zonabris floralis, M. fuesslini Panzer 1793, Zonabris fuesslini] G-PR05

Hycleus scutellatus (Rosenhauer 1856) [=Mylabris scutellata, Zonabris scutellata; incl. Zonabris decempunctata var. arapilensis Redondo 1915, Z. (Hicleus) decempunctata, Z. decimpunctata var. deficiens De la Fuente 1910, Mylabris lichtensteini Górriz Muñoz 1882, Z. decempunctata var. salmanticensis Redondo 1915] G-PR05

Hycleus wagneri (Chevrolat 1837) [=Mylabris wagneri; incl. M. curta Chevrolat 1837, Zonabris insignipennis Pic 1901] G-PR05

Lagorina sericea (Waltl 1835) [=Cantharis sericea, Lytta (Lagorina) sericea; incl. L. herbivora Dej. in Amor Mayor 1860 (n. n.)] G-PR05

Lytta vesicatoria (Linnaeus 1758) [=Meloe vesicatorius, Cantharis vesicatoria, Litta (l. c.) vesicatoria; incl. Lytta vesicatoria var. aurantiaca Escherich 1894] G-PR05

Nemognatha chrysomelina (Fabricius 1775) [=Nemognata (l. c.) chrysomelina; incl. Nemognatha chrysomelina ab. nigripes Sufrian 1853] G-PR05

Sitaris muralis (Foerster 1771) G-PR05 [incl. Si. humeralis Westwood 1839 G-PR05, Necydalis humeralis L02, Sytaris (l. c.) humeralis G-PR05, Si. muralis var. splendida Schaufuss 1861 G-PR05]

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[BB68] Basford, N. L., J. E. Butler, C. A. Leone & F. J. Rohlf. 1968. Immunologic comparisons of selected Coleoptera with analyses of relationships using numerical taxonomic methods. Systematic Zoology 17 (4): 388–406.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C96] Campbell, N. A. 1996. Biology (4th ed.) The Benjamin/Cummings Publishing Company, Inc.: Menlo Park (California).

[C90] Crawford, C. S. 1990. Scorpiones, Solifugae, and associated desert taxa. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 421–475. John Wiley & Sones: New York.

[C01a] Csiki, E. 1901a. Utivázlat [Reiseskizze]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. xii–xli. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[C01b] Csiki, E. 1901b. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers vol. 1. Melbourne University Press: Carlton (Victoria).

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G-PP-OS05] García-París, M., G. Parra-Olea & C. Settanni. 2005. Primer registro de Epicauta segmenta (Say, 1824) (Coleoptera, Meloidae) en Querétaro (México). Graellsia 61 (2): 257–258.

[G-PR05] García-París, M., & J. L. Ruiz. 2005. Bibliografía y registros Ibero-Baleares de Meloidae (Coleoptera) publicados hasta la aparición del “Catálogo sistemático geográfico de los coleópteros observados en la Península Ibérica, Pirineos propiamente dichos y Baleares” de J. M. de la Fuente (1933). Graellsia 61 (2): 225–255.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[R35] Rayment, T. 1935. A Cluster of Bees: Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species. Endeavour Press: Sydney.

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