Dorsum of Trionyxella granulata, from Roewer (1935).

Belongs within: Assamiidae.

The Trionyxellinae are a group of harvestmen found in southern Africa and Asia, characterised by the presence of a pseudonychium between the true claws.

Characters (from Roewer 1935): Anterior margin of the carapace unarmed medially; spiracles concealed by chitinous projections; femur of pedipalp with short ventral teeth; distitarsus I two-segmented; tarsi III and IV with simple claws, with pseudonychium present over claws.

<==Trionyxellinae [Trionyxellidae]
    |--Triaenopodium Roewer 1915 R35
    |    `--T. tarsalis Roewer 1915 R35
    |--Kandyca Roewer 1915 R35
    |    `--K. minima Roewer 1915 R35
    |--Nuwaria Roewer 1915 R35
    |    `--N. granulata Roewer 1915 R35
    |--Nilgirius Roewer 1915 R35
    |    `--N. scaber Roewer 1915 R35
    |--Calloristus Roewer 1935 R35
    |    `--*C. granipes Roewer 1935 R35
    |--Balnissa Roewer 1935 R35
    |    `--*B. parva Roewer 1935 R35
    |--Brysma Roewer 1935 R35
    |    `--*B. atra Roewer 1935 R35
    |--Trionychiperna Roewer 1929 R35
    |    `--T. carli Roewer 1929 R35
    |--Sampitius Roewer 1935 R35
    |    `--*S. opacus Roewer 1935 R35
    |--Dulitellus Roewer 1935 R35
    |    |--*D. maculatus Roewer 1935 R35
    |    `--D. sarawakensis WP-G10
    |--Namutonia Lawrence 1931 R35 [incl. Gobabisia Roewer 1940 S92]
    |    |--*N. scabra Lawrence 1931 S92
    |    `--N. wuehlischi (Roewer 1940) [=Gobabisia wuehlischi] S92
    |--Trionyxella Roewer 1912 R35
    |    |--T. clavipus Roewer 1912 R35
    |    `--T. granulata Roewer 1935 R35
    `--Randilella Lawrence 1963 L63
         `--*R. transvaalensis Lawrence 1963 S92

*Type species of generic name indicated


[L63] Lawrence, R. F. 1963. The Opiliones of the Transvaal. Annals of the Transvaal Museum 24 (4): 275–304.

[R35] Roewer, C.-F. 1935. Alte und neue Assamiidae. Weitere Weberknechte VIII. (8. Ergänzung der “Weberknechte der Erde” 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Uebersee-Museum in Bremen 1: 1–168.

[S92] Staręga, W. 1992. An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum 33 (2): 271–336.

[WP-G10] Willemart, R. H., A. Pérez-González, J.-P. Farine & P. Gnaspini. 2010. Sexually dimorphic tegumental gland openings in Laniatores (Arachnida, Opiliones), with new data on 23 species. Journal of Morphology 271 (6): 641–653.

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