Staphylinini

Acylophorus pratensis, copyright Richard Migneault.

Belongs within: Staphylininae.
Contains: Amblyopinina, Cyrtoquediina, Indoquedius, Quedius, Philonthina, Staphylinina.

The Staphylinini are a cosmopolitan group of relatively large staphylinid beetles, including the rove beetles that are the most familiar members of the family.

Staphylinini [Quediini] N90
    |  i. s.: Valdiviodes BC16
    |         Alesiella BC16
    |         Quediomacrus BC16
    |         Lonia BC16
    |         Quelaestrygon BC16
    |         Strouhalium gracilicorne BC16
    |--Cretoquedius BC16
    `--+--Atanygnathus Jacobson 1909 NT01 (see below for synonymy)
       |    `--A. acuminatus BC16
       `--+--Antimerus punctipennis BC16
          |--+--Afroquedius sexpunctatus BC16
          |  `--Amblyopinina BC16
          `--+--Cyrtoquediina BC16
             `--+--+--Erichsonius Fauvel 1874 NT01 (see below for synonymy)
                |  |    |--E. nanus BC16
                |  |    `--E. patella N90
                |  `--Acylophorina BC16
                |       |  i. s.: Australotarsius grandis BC16
                |       |         Acylohsellus BC16
                |       |         Stevensia longipennis BC16
                |       |--Hemiquedius Casey 1915 BC16, NT01
                |       |    `--H. ferox (LeConte 1878) NT01
                |       `--+--Anaquedius Casey 1915 BC16, NT01
                |          |    `--A. vernix (LeConte 1878) NT01
                |          `--+--+--Anchocerus BC16
                |             |  `--Paratolmerus siamensis BC16
                |             `--Acylophorus Nordmann 1837 BC16, NT01 (see below for synonymy)
                |                  |--A. capensis BC16
                |                  |--A. pratensis BC16
                |                  `--A. wagenschieberi BC16
                `--+--Indoquedius BC16
                   `--+--Quediina BC16
                      |    |--Quedius BC16
                      |    |--Pseudorientis BC16
                      |    |--Velleiopsis BC16
                      |    `--Beeria Hatch 1957 NT01
                      |         `--B. nematocera (Casey 1915) [incl. B. punctata Hatch 1957] NT01
                      `--Staphylinini propria BC16
                           |  i. s.: Eucibdelus BC16
                           |         Ioma BC16
                           |         Weiserianum BC16
                           |         Haematodes BC16
                           |         Euristus BC16
                           |--+--Philothalpus BC16
                           |  |    |--P. bilobus BC16
                           |  |    `--P. falini BC16
                           |  `--Xanthopygina BC16
                           |       |  i. s.: Gastrisus Sharp 1876 NT01
                           |       |         Trigonopselaphus Gemminger & Harold 1868 (see below for synonymy) NT01
                           |       |--+--Isanopus sallaei BC16
                           |       |  `--+--Nordus fungicola BC16
                           |       |     `--Plociopterus BC16
                           |       `--+--Glenus flohri BC16
                           |          `--Xanthopygus Kraatz 1857 BC16, GN09 (see below for synonymy)
                           |               |--X. chapareanus BC16
                           |               `--X. xanthopygus Nordmann 1837 [incl. X. cacti Horn 1868] NT01
                           `--+--+--Philonthina BC16
                              |  `--Algon BC16
                              |       |--A. hollowayae BC16
                              |       `--A. oculatus BC16
                              `--+--Staphylinina BC16
                                 `--Anisolinina BC16
                                      |--+--Misantlius gebieni BC16
                                      |  `--Hesperosoma pederseni BC16
                                      `--+--Tolmerinus BC16
                                         `--+--Tympanophorus Nordmann 1837 BC16, NT01
                                            |    |--T. concolor Sharp 1884 NT01
                                            |    `--T. puncticollis Erichson 1840 (see below for synonymy) NT01
                                            `--+--Anisolinus BC16
                                               `--Pammegus ruficollis BC16

Acylophorus Nordmann 1837 BC16, NT01 [incl. Amacylophorus Smetana 1971 NT01, Palpacylophorus Smetana 1971 NT01]

Atanygnathus Jacobson 1909 NT01 [incl. Tanygnathinus Reitter 1909 NT01, Tanygnathus Erichson 1839 non Wagler 1832 NT01; Tanygnathinina BC16]

Erichsonius Fauvel 1874 NT01 [incl. Actobius Fauvel 1876 NT01, Sectophilonthus Tottenham 1949 NT01; Erichsoniina BC16]

Trigonopselaphus Gemminger & Harold 1868 [incl. Trigonophorus Nordmann 1837 nec Stephens 1829 nec Solier 1851] NT01

Tympanophorus puncticollis Erichson 1840 [incl. Xanthopygus borealis Hatch 1957, T. borealis] NT01

Xanthopygus Kraatz 1857 BC16, GN09 [incl. Heteropygus Bernhauer 1906 NT01, Lampropygus Sharp 1884 NT01]

*Type species of generic name indicated

REFERENCES

[BC16] Brunke, A. J., S. Chatzimanolis, H. Schillhammer & A. Solodovnikov. 2016. Early evolution of the hyperdiverse rove beetle tribe Staphylinini (Coleoptera: Staphylinidae: Staphylininae) and a revision of its higher classification. Cladistics 32 (4): 427–451.

[GN09] Grebennikov, V. V., & A. F. Newton. 2009. Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera). European Journal of Entomology 106: 275–301.

[N90] Newton, A. F., Jr. 1990. Insecta: Coleoptera Staphylinidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1137–1174. John Wiley & Sones: New York.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.