Gabronthus thermarum, copyright Naturalis, Biodiversity Centre.

Belongs within: Staphylininae.
Contains: Philonthus.

The Staphylinini are a cosmopolitan group of relatively large staphylinid beetles, including the rove beetles that are the most familiar members of the family.

Staphylinini N90
    |--Amblyopinina [Amblyopinini] B14
    |    |--Myotyphlus jansoni A71, LB91
    |    |--Edrabius A71
    |    |--Megamblyopinus A71
    |    |--Amblyopinus A71
    |    `--Amblyopinodes piceus (Brèthes 1926) B14
    `--Philonthina B14
         |--Actinus B14
         |    |--A. imperialis Fauvel 1878 B14
         |    `--A. macleayi B14
         |--Philonthus N90
         |--Gabronthus Tottenham 1955 T55
         |    |--*G. maritimus [=Gabrius maritimus, Philonthus maritimus] T55
         |    |--G. alluaudanus (Jeannel & Paulian 1945) [=Gabrius alluaudanus] T55
         |    |--G. badalus Tottenham 1955 T55
         |    |--G. chyuluensis (Cameron 1942) [=Philonthus chyuluensis] T55
         |    |--G. corrugatus Tottenham 1955 T55
         |    |--G. fundator Tottenham 1955 T55
         |    |--G. gombanus Tottenham 1955 T55
         |    |--G. inclinans (Walker 1859) [=Philonthus inclinans; incl. P. angustatus] T55
         |    |--G. kisiranus Tottenham 1955 T55
         |    |--G. kosanus Tottenham 1955 T55
         |    |--G. meridionalis [=Philonthus meridionalis] T55
         |    |--G. mgogorius Tottenham 1955 T55
         |    |--G. minusculus Tottenham 1955 T55
         |    |--G. minutissimus (Bernahuer 1915) [=Philonthus minutissimus] T55
         |    |--G. oribates (Jeannel & Paulian 1945) [=Gabrius oribates] T55
         |    |--G. pluvialis Tottenham 1955 T55
         |    |--G. pygmaeus (Kraatz 1859) [=Philonthus pygmaeus] T55
         |    |--G. styx Tottenham 1955 T55
         |    |--G. sulcifrons (Sharp 1889) [=Philonthus sulcifrons] T55
         |    |--G. tenuissimus (Tottenham 1949) [=Philonthus tenuissimus] T55
         |    |--G. thermaroides Tottenham 1955 T55
         |    |--G. thermarum (Aubé 1850) [=Philonthus thermarum, Gabrius thermarum] T55
         |    `--G. turneri (Tottenham 1949) [=Philonthus turneri] T55
         `--Gabrius T55
              |--*G. politus [=Philonthus politus] T55
              |--G. kamatembeanus [=Philonthus (Gabrius) kamatembeanus] T55
              |--G. separatus [=Philonthus (Gabrius) separatus] T55
              `--G. ‘turneri’ (Cameron 1950) [=Philonthus (Gabrius) turneri non P. turneri Tottenham 1949] T55

Staphylinini incertae sedis:
  Tympanophorus puncticollis N90
  Erichsonius patella N90
  Neobisnius N90
  Belonuchus penetrans Silvestri 1945 N90, S57
  Hesperus baltimorensis N90
  Platydracus Thomson 1858 N90, GN09
    |--P. maculosus (Gravenhorst 1802) GN09
    |--P. viridanus N90
    `--P. zonatus N90
  Staphylinus Linnaeus 1758 L58
    |--S. arenarius L02
    |--S. badipes N90
    |--S. boleti Linnaeus 1758 L58
    |--S. canaliculatus [=Aleochara canaliculata] L02
    |--S. caraboides Linnaeus 1758 L58
    |--S. chrysomelinus Linnaeus 1758 L58
    |--S. dauricus C01
    |--S. erytropterus Linnaeus 1758 L58
    |--S. flavescens Linnaeus 1758 L58
    |--S. flavipes Linnaeus 1758 L58
    |--S. fulgidus [=Paederus fulgidus] L02
    |--S. fuscipes Linnaeus 1758 L58
    |--S. hirtus Linnaeus 1758 L58
    |--S. lignorum Linnaeus 1758 L58
    |--S. maxillosus Linnaeus 1758 L58
    |--S. murinus Linnaeus 1758 L58
    |--S. olens L02
    |--S. picipennis C01
    |--S. plagiatus L02
    |--S. politus Linnaeus 1758 L58
    |--S. riparius Linnaeus 1758 L58
    |--S. rufipes Linnaeus 1758 L58
    |--S. rufus Linnaeus 1758 L58
    |--S. sanguineus Linnaeus 1758 L58
    |--S. similis C01
    |--S. stercorarius C01
    `--S. subterraneus Linnaeus 1758 L58

*Type species of generic name indicated


[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[GN09] Grebennikov, V. V., & A. F. Newton. 2009. Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera). European Journal of Entomology 106: 275–301.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[N90] Newton, A. F., Jr. 1990. Insecta: Coleoptera Staphylinidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1137–1174. John Wiley & Sones: New York.

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

[T55] Tottenham, C. E. 1955. Studies in the genus Philonthus Stephens (Coleoptera: Staphylinidae). Parts II, III, and IV. Transactions of the Royal Entomological Society of London 106 (3): 153–195.

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