Harpalus affinis, copyright Trevor & Dilys Pendleton.

Belongs within: Harpalitae.

Harpalus is a diverse genus of carabid beetles with antennae that are setose from the third segment onwards, and two ventral rows of scale-like setae on the fore tarsi of males. They are native to Eurasia and North America though some species have also been introduced outside this range.

Harpalus Latreille 1802 LM87
    |--H. affinis (Schrank 1781) E04, LM87 (see below for synonymy)
    |--H. aeneus C01
    |--H. amariformis C01
    |--H. ambiguus [=Carabus ambiguus] L02
    |--H. caerulescens (Linnaeus 1758) L02, L58 [=Carabus caerulescens L02]
    |--H. caliginosus BB68
    |--H. calydonius [=Carabus calydonius] L02
    |--H. cephalotes (Linnaeus 1758) L02, L58 [=Carabus cephalotes L02]
    |--H. cisteloides [=Carabus cisteloides] L02
    |--H. cupreus (Linnaeus 1758) L02, L58 [=Carabus cupreus L02]
    |--H. distinguendus C01
    |--H. eraticus BS02
    |--H. fulvicornis (Thunberg 1806) [=Carabus fulvicornis] LM87
    |--H. fuscicalpis C01
    |--H. gagates [=Scarites gagates] L02
    |--H. germanus (Linnaeus 1758) L02, L58 [=Carabus germanus L02]
    |--H. gory Gory 1833 LM87
    |--H. goryi Boisduval 1835 LM87
    |--H. ignobilis Boheman 1858 LM87
    |--H. indianus BS02
    |--H. interruptus [=Carabus interruptus] L02
    |--H. leucopthalmus [=Carabus leucopthalmus] L02
    |--H. marginatus (Linnaeus 1758) L02, L58 [=Carabus marginatus L02]
    |--H. marginepunctatus B35
    |--H. metallicus [=Carabus metallicus] L02
    |--H. motschulskyi C01
    |--H. pennsylvanicus BB68
    |--H. piceus [=Scarites piceus] L02
    |--H. picipennis C01
    |--H. prasinus [=Carabus prasinus] L02
    |--H. puncticeps E04
    |--H. rubripes (De Geer 1774) VPB11
    |--H. rufibarbis E04
    |--H. rufipes (De Geer 1774) VPB11 [=Cucujus rufipes L02]
    |--H. serripes C01
    |--H. sexpunctatus (Linnaeus 1758) L02, L58 [=Carabus sexpunctatus L02]
    |--H. spiniger [=Carabus spiniger] L02
    |--H. striatus [=Carabus striatus] L02
    |--H. striola [=Carabus striola] L02
    |--H. tardus (Panzer 1796) VPB11
    |--H. terricola [=Carabus terricola] L02
    |--H. vestitus [=Carabus vestitus] L02
    `--H. vulgaris (Linnaeus 1758) L02, L58 [=Carabus vulgaris L02]

Harpalus affinis (Schrank 1781) E04, LM87 [=Carabus affinis LM87; incl. C. proteus Paykull 1790 LM87, *Harpalus proteus LM87]

*Type species of generic name indicated


[BS02] Ball, G. E., & D. Shpeley. 2002. Ginemini, Ginema thomasi, new tribe, new genus and new species, from Amazonian Bolivia (Coleoptera: Carabidae: Harpalinae). Transactions of the American Entomological Society 128 (1): 75–98.

[BB68] Basford, N. L., J. E. Butler, C. A. Leone & F. J. Rohlf. 1968. Immunologic comparisons of selected Coleoptera with analyses of relationships using numerical taxonomic methods. Systematic Zoology 17 (4): 388–406.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E04] Emberson, R. M. 2004. A European ground beetle, Harpalus tardus (Panzer) (Coleoptera: Carabidae), in Canterbury, and the predisposition of European carabids to establish in exotic temperate locations. New Zealand Entomologist 27: 39–42.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

Last updated: 5 March 2018.

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