Harpalus affinis, copyright Trevor & Dilys Pendleton.

Belongs within: Harpalinae.

Harpalus is a diverse genus of carabid beetles with antennae that are setose from the third segment onwards, and two ventral rows of scale-like setae on the fore tarsi of males.

Harpalus Latreille 1802 [Harpalina] LM87
    |--H. affinis (Schrank 1781) VPB11, LM87 (see below for synonymy)
    |--H. aeneus C01
    |--H. amariformis C01
    |--H. ambiguus [=Carabus ambiguus] L02
    |--H. caerulescens (Linnaeus 1758) L02, L58 [=Carabus caerulescens L02]
    |--H. caliginosus BB68
    |--H. calydonius [=Carabus calydonius] L02
    |--H. cephalotes (Linnaeus 1758) L02, L58 [=Carabus cephalotes L02]
    |--H. cisteloides [=Carabus cisteloides] L02
    |--H. cupreus (Linnaeus 1758) L02, L58 [=Carabus cupreus L02]
    |--H. distinguendus C01
    |--H. eraticus BS02
    |--H. fulvicornis Thunberg 1806) [=Carabus fulvicornis] LM87
    |--H. fuscicalpis C01
    |--H. gagates [=Scarites gagates] L02
    |--H. germanus (Linnaeus 1758) L02, L58 [=Carabus germanus L02]
    |--H. gory Gory 1833 LM87
    |--H. goryi Boisduval 1835 LM87
    |--H. ignobilis Boheman 1858 LM87
    |--H. indianus BS02
    |--H. interruptus [=Carabus interruptus] L02
    |--H. leucopthalmus [=Carabus leucopthalmus] L02
    |--H. marginatus (Linnaeus 1758) L02, L58 [=Carabus marginatus L02]
    |--H. metallicus [=Carabus metallicus] L02
    |--H. motschulskyi C01
    |--H. pennsylvanicus BB68
    |--H. piceus [=Scarites piceus] L02
    |--H. picipennis C01
    |--H. prasinus [=Carabus prasinus] L02
    |--H. puncticeps E04
    |--H. rubripes (De Geer 1774) VPB11
    |--H. rufibarbis E04
    |--H. rufipes (De Geer 1774) VPB11 [=Cucujus rufipes L02]
    |--H. serripes C01
    |--H. sexpunctatus (Linnaeus 1758) L02, L58 [=Carabus sexpunctatus L02]
    |--H. spiniger [=Carabus spiniger] L02
    |--H. striatus [=Carabus striatus] L02
    |--H. striola [=Carabus striola] L02
    |--H. tardus (Panzer 1796) VPB11
    |--H. terricola [=Carabus terricola] L02
    |--H. vestitus [=Carabus vestitus] L02
    `--H. vulgaris (Linnaeus 1758) L02, L58 [=Carabus vulgaris L02]

Harpalus affinis (Schrank 1781) VPB11, LM87 [=Carabus affinis LM87; incl. C. proteus Paykull 1790 LM87, *Harpalus proteus LM87]

*Type species of generic name indicated


[BS02] Ball, G. E., & D. Shpeley. 2002. Ginemini, Ginema thomasi, new tribe, new genus and new species, from Amazonian Bolivia (Coleoptera: Carabidae: Harpalinae). Transactions of the American Entomological Society 128 (1): 75–98.

[BB68] Basford, N. L., J. E. Butler, C. A. Leone & F. J. Rohlf. 1968. Immunologic comparisons of selected Coleoptera with analyses of relationships using numerical taxonomic methods. Systematic Zoology 17 (4): 388–406.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E04] Emberson, R. M. 2004. A European ground beetle, Harpalus tardus (Panzer) (Coleoptera: Carabidae), in Canterbury, and the predisposition of European carabids to establish in exotic temperate locations. New Zealand Entomologist 27: 39–42.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

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