Bertkauia crosbyana, copyright Tom Murray.

Belongs within: Epipsocetae.
Contains: Epipsocus.

The Epipsocidae are a group of bark-lice characterised by the presence of a pair of sclerotised ridges running the length of the labrum. Subfamilies have been recognised on the basis of fore wing venation, with species of Gojinae having more than three branches each in vein Rs and M whereas Epipsocinae have two branches to Rs and three to M (Li 2002), but the significance of this distinction is questionable (New & Liernhard 2007).

Characters (from New & Lienhard 2007): Fore wing length usually 2.5–5.7 mm. Head elongate. Labrum with two sclerotised rods/ridges running entire length of labrum, reaching sides of labrum anteriorly and joined, if at all, by only a weakly sclerotised anterior connection. Antennal scape membranous over much of ventral surface. Tarsi 2-segmented, claws with or without subapical tooth. Anterior and hind femora sometimes with row of cones at bases of setae. Macropterous or brachypterous, females rarely apterous. Fore wing pterostigma without crossveins; areola postica free; macropterous individuals with one anal vein in fore wing. Female with ventral gonapophysis valve present or absent; external valve reduced, represented as small swelling or setal field on dorsal valve.

    |  i. s.: Mesepipsocus mobilis (Hagen 1861) (see below for synonymy) M74
    |--Gojinae NL07
    |    |--Goja NL07
    |    `--Cubitiglabra Li 1995 L03, L02 [=Cubitiglabrus (l. c.) L03, Cubitiglatrus (l. c.) L03]
    |         |--*C. quadripunctatus Li 1995 L02
    |         |--C. dayaoshanana Li 2002 [=Cubitiglabrus (l. c.) dayaoshananus] L03
    |         `--C. polyphlebius Li 1995 L02
    `--Epipsocinae NL07
         |--Epipsocus NL07
         |--Liratepipsocus Li 2002 L02
         |    `--*L. jinghongicus Li 2002 L02
         |--Metepipsocus Li 2002 L02
         |    `--*M. beijingicus Li 2002 L02
         |--Valvepipsocus Li 2002 L02 [=Valviepipsocus L03]
         |    `--*V. diodematus Li 2002 L03
         |--Hinduipsocus Badonnel 1981 L02
         |    |--*H. annulipes Badonnel 1981 L02
         |    `--H. hongkongensis Li & Mockford 1997 L02
         |--Dichoepipsocus Li & Mockford 1997 L02
         |    |--*D. micropterus Li & Mockford 1997 L02
         |    `--D. dictyodromus Li 2002 L02
         |--Spordoepipsocus Li 2002 L02
         |    |--*S. perforatus Li 2002 L02
         |    |--S. formosus (Li 1992) [=Epipsocus formosus] L02
         |    `--S. imperforatus Li 2002 L02
         |--Dicropsocus NL07
         |    |--*D. montanus Smithers & Thornton 1977 NL07
         |    |--D. complexus Smithers & Thornton 1977 NL07
         |    `--D. rugosus Smithers & Thornton 1977 NL07
         |--Heteroepipsocus Li 1995 L02
         |    |--*H. longicellus Li 1995 L02
         |    |--H. brevicellus Li 1995 L02
         |    |--H. inornatus (Banks 1916) [=Epipsocus inornatus] NL07
         |    `--H. maculatus Li 2002 L02
         |--Bertkauia Kolbe 1882 L02 [=Bertkallis (l. c.) L03; incl. Lapithes Bertkau 1883 L02]
         |    |--B. lucifuga (Rambur 1842) M93, G74 (see below for synonymy)
         |    |--B. crosbyana Chapman 1930 [=Epipsocus crosbyanus] M93
         |    |--B. lepicidinaria Chapman 1930 [=Epipsocus lepicidinarius] M93
         |    |--B. loebli M03
         |    |--B. marusiki Mockford 2003 M03
         |    |--B. minuscula Mockford 2003 M03
         |    |--B. remyi M03
         |    `--B. reticularis Li & Mockford 1997 L02
         `--Epipsocopsis Badonnel 1955 L02 [incl. Epipsocidus Badonnel 1955 NL07, B55]
              |--*E. machadoi Badonnel 1955 L02
              |--E. angolensis (Badonnel 1955) NL07, B55 [=Epipsocus (*Epipsocidus) angolensis NL07]
              |--E. apicalis New & Thornton 1976 NL07
              |--E. basalis New & Thornton 1976 NL07
              |--E. cameronensis New & Lee 1991 NL07
              |--E. completa (Banks 1916) NL07
              |--E. corollifer Li 2002 L02
              |--E. costalis (Banks 1914) NL07
              |--E. dubia (Karny 1925) NL07
              |--E. fasciata Smithers & Thornton 1977 NL07
              |--E. fumipennis (Banks 1920) NL07
              |--E. hyalina (Banks 1920) NL07
              |--E. longiceps (Enderlein 1926) NL07
              |--E. maclurei New & Thornton 1976 NL07
              |--E. macrostigma Smithers & Thornton 1977 NL07
              |--E. maculata New & Thornton 1976 NL07
              |--E. magna (New & Thornton 1976) NL07
              |--E. mouldsi Smithers 1976 NL07
              |--E. murca (Enderlein 1903) NL07
              |--E. murphyi New & Thornton 1976 NL07
              |--E. nebulifera Smithers & Thornton 1977 NL07
              |--E. nubilipennis (Karny 1925) NL07
              |--E. paraselena Vaughan, Thorton & New 1989 NL07
              |--E. peradenayense New 1977 L02
              |--E. prominens (Banks 1937) NL07
              |--E. punctata Smithers & Thornton 1977 NL07
              |--E. sclerota New & Thornton 1976 NL07
              |--E. selena New & Thornton 1976 NL07
              |--E. serpentinus Li 2002 L02
              |--E. setosa New & Thornton 1976 NL07
              |--E. singaporensis New & Thornton 1976 NL07
              |--E. thailandensis New 1973 NL07
              `--E. vilhenai Badonnel 1955 B55

Bertkauia lucifuga (Rambur 1842) M93, G74 [=Psocus lucifugus G74, Epipsocus lucifugus G74; incl. *Bertkauia prisca Kolbe 1882 NL07, G74, Lapithes pulicarius Bertkau 1883 L98]

Mesepipsocus mobilis (Hagen 1861) [=Psocus mobilis, Caecilius mobilis, Epipsocus mobilis, Psocidus mobilis; incl. Mesepipsocus grassei Badonnel 1969] M74

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103–215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[M03] Mockford, E. L. 2003. New species and records of Psocoptera from the Kuril Islands. Mitt. Mus. Nat.kd. Berl., Dtsch. Entomol. Z. 50 (2): 191–230.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

Last updated: 4 August 2020.

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