Bembidiinae

Micratopus sp., copyright Mike Quinn.


Belongs within: Trechinae.
Contains: Bembidiina, Tachyina.

The Bembidiinae are a group of carabid beetles with reduced terminal segments on the labial and maxillary palps, and two unequally dilated segments in the tarsi of males.

Bembidiinae
    |--Micratopus B08 [Micratopini B90]
    `--Bembidiini R-JC01
         |  i. s.: Anillinus B90
         |         Phrypeus rickseckeri B90
         |         Asaphidion B90
         |           |--A. cyanicorne F05
         |           |    |--A. c. cyanicorne F05
         |           |    `--A. c. atlanticum Ortuño & Toribio 2005 F05
         |           |--A. flavipes (Linnaeus 1761) [=Cicindela flavipes] B55
         |           `--A. stierlini (Heyden 1880) VPB11
         |         Lymneops B90
         |         Mioptachys flavicauda (Say 1823) B90, B14
         |         Tachyta Kirby 1837 B90, LM87
         |           |  i. s.: T. subvirens B55
         |           `--T. (Tachyta) LM87
         |                |--T. (T.) nana LM87
         |                |    |--T. n. nana LM87
         |                |    `--T. n. inornatus (Say 1823) [incl. *T. picipes Kirby 1837] LM87
         |                `--T. (T.) brunnipennis (Macleay 1871) [=Bembidium brunnipenne] LM87
         |         Porotachys bisulcatus B90
         |         Illaphanus Macleay 1865 LB91, LM87
         |           |--*I. stephensii Macleay 1865 LM87
         |           `--I. macleayi Lea 1906 LM87
         |--Bembidiina R-JC01
         |--Tachyina R-JC01
         `--Gouleta [Xystosomina] R-JC01
              `--G. cayennensis R-JC01

Bembidiinae incertae sedis:
  Lymnastis Motschulsky 1862 B08
    |--*L. indicus (Motschulsky 1851) [=Lymnaeum indicum] B08
    |--L. atricapillus B08
    |--L. brooksi Baehr 2008 B08
    |--L. coomani B08
    |--L. decorsei B55
    |--L. inops B08
    |--L. macrops B55
    |--L. niloticus B55
    |--L. pilosus Bates 1892 [incl. Tachys setiger Sloane 1903] LM87
    `--L. pullulus B08
  Brachytachys Basilewsky 1953 B55
    |--B. curtulus Basilewsky 1953 B55
    `--B. villiersi Basilewsky 1953 B55
  Sphaerotachys B55
    |--S. fumigatus B55
    `--S. haemorrhoidalis B55
  Tachyura B55
    |--T. apicalis B55
    |--T. conspicua B55
    |--T. emerita B55
    |--T. fausta B55
    |--T. grandicollis [=Amaurotachys grandicollis] B55
    |    |--T. g. grandicollis B55
    |    |--T. g. michaelseni B55
    |    `--T. g. nigrolimbatus B55
    |--T. vagans B55
    |--T. variabilis B55
    `--T. viverra B55

*Type species of generic name indicated

REFERENCES

[B08] Baehr, M. 2008. A new cavernicolous and eyeless species of the genus Lymnastis from Christmas Island, Australia (Insecta: Coleoptera: Carabidae: Bembidiinae). Records of the Western Australian Museum 24 (2): 199–203.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

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