Trimalaconothrus

Dorsal (left) and ventral views of Trimalaconothrus platyrhinus, from Balogh & Balogh (1992, vol. 2).

Belongs within: Desmonomata.

Trimalaconothrus is a cosmopolitan genus of oribatid mites characterised by a covering of birefringent cerotegument together with tridactyl legs.

Characters (from Balogh & Balogh 1992, as Trimalaconothrus + Zeanothrus + Fossonothrus): Bothridial sensillus absent. Cerotegument birefringent, porose. Opisthosomal gland absent. Epimeral setal formula 3-1-3-3 or 3-2-3-3. Genital and anal plates of macropyline type, large, occupying entire ventral side posterior to epimeres; four to twelve pairs of genital setae; aggenital setae absent; one pair of anal setae; three pairs of adanal setae. Legs with three tarsal claws.

<==Trimalaconothrus Berlese 1916 S04
    |--T. (Trimalaconothrus) [incl. Zeanothrus Hammer 1966] S04
    |    |--*T. (T.) indusiatus (Berlese 1916) [=Malaconothrus (*T.) indusiatus] S04
    |    |--T. (T.) africanus Balogh 1958 (n. d.) S04
    |    |--T. (T.) almagrensis Iglesias, Palacios-Vargas & Mahunka 2001 S04
    |    |--T. (T.) altissimus Piffl 1971 S04
    |    |--T. (T.) angulatus Willmann 1931 S04
    |    |--T. (T.) aureopunctatus (Hammer 1979) [=Malaconothrus aureopunctatus] S04
    |    |--T. (T.) azumaensis Yamamoto, Kuriki & Aoki 1993 S04
    |    |--T. (T.) barbatus Yamamoto 1977 S04
    |    |--T. (T.) brevisetiger Yamamoto & Aoki 1998 S04
    |    |--T. (T.) crinitus (Berlese 1908) [=Malaconothrus crinitus] S04
    |    |--T. (T.) crispus Hammer 1962 S04
    |    |--T. (T.) eichhornicus Iglesias, Palacios-Vargas & Mahunka 2001 S04
    |    |--T. (T.) elegans (Hammer 1966) S04 [=*Zeanothrus elegans BB92]
    |    |--T. (T.) flagelliformis Wallwork 1970 S04
    |    |--T. (T.) foveolatus Willmann 1931 S04
    |    |    |--T. f. foveolatus S04
    |    |    `--T. f. sundaicus Willmann 1931 S04
    |    |--T. (T.) grandis Hammen 1952 S04
    |    |--T. (T.) granulatus Tseng 1982 S04
    |    |--T. (T.) hakonensis Yamamoto 1977 S04
    |    |--T. (T.) heterotrichus Wallwork 1973 S04
    |    |--T. (T.) iteratus Subías 2004 [=T. longirostris Hammer 1966 non Willmann 1929] S04
    |    |--T. (T.) joonsooi Choi 1986 S04
    |    |--T. (T.) kurikii Yamamoto 1977 S04
    |    |--T. (T.) lineolatus Balogh & Balogh 1986 S04
    |    |--T. (T.) longirostris Willmann 1929 S04
    |    |--T. (T.) longisetosus Yamamoto & Aoki 2000 S04
    |    |--T. (T.) maniculatus Fain & Lambrechts 1987 S04
    |    |--T. (T.) mirabilis Sergienko & Melamud 1993 S04
    |    |--T. (T.) montanus Hammer 1958 S04
    |    |--T. (T.) multipilosus Balogh & Balogh 2002 S04
    |    |--T. (T.) nipponicus Yamamoto & Aoki 1971 S04
    |    |--T. (T.) nodosus Yamamoto 1997 S04
    |    |--T. (T.) obsessus Subías 2004 [=T. albulus Tseng 1982 non Hammer 1972] S04
    |    |--T. (T.) opisthoseta Hammer 1966 S04
    |    |--T. (T.) oppositus Balogh 1997 S04
    |    |--T. (T.) oxyrhinus Hammer 1962 S04
    |    |--T. (T.) pallidus Wallwork 1977 S04
    |    |--T. (T.) pilipes Willmann 1933 S04
    |    |--T. (T.) planus Balogh 1958 (n. d.) S04
    |    |--T. (T.) platyrhinus Hammer 1962 S04
    |    |--T. (T.) prahuensis Hammer 1979 S04
    |    |--T. (T.) rafalskii Olszanowski 1997 S04
    |    |--T. (T.) repetitus Subías 2004 [=T. reticulatus Yamamoto 1977 non Balogh 1958] S04
    |    |--T. (T.) reticulatus Balogh 1958 (n. d.) S04
    |    |--T. (T.) sacculus Hammer 1966 S04
    |    |--T. (T.) scimitarum Tarras-Wahlberg 1985 S04
    |    |--T. (T.) scutatus Mihelčič 1959 S04
    |    |--T. (T.) similis Balogh 1958 (n. d.) S04
    |    |--T. (T.) simplex (Banks 1895) [=Nothrus simplex] S04
    |    |--T. (T.) soror Balogh 1958 (n. d.) S04
    |    |--T. (T.) tardus (Michael 1888) [=Nothrus tardus] S04
    |    |--T. (T.) tonkini Luxton 1982 S04
    |    |--T. (T.) vietsi Willmann 1925 S04
    |    |--T. (T.) wallworki (Starý & Block 1995) [=Fossonothrus wallworki] S04
    |    `--T. (T.) yachidairaensis Yamamoto, Kuriki & Aoki 1993 S04
    `--T. (Tyrphonothrus Knülle 1957) [incl. Fossonothrus Hammer 1962] S04
         |--T. (T.) maior (Berlese 1910) (see below for synonymy) S04
         |--T. (T.) albulus Hammer 1972 S04
         |--T. (T.) angustirostrum Hammer 1966 S04
         |--T. (T.) australis Hammer 1958 S04
         |--T. (T.) barrancensis Hammer 1961 S04
         |--T. (T.) blancus Hammer 1961 S04
         |--T. (T.) buresi Kunst 1959 S04
         |--T. (T.) cajamarcensis Hammer 1961 S04
         |--T. (T.) canopeus Iglesias, Palacios-Vargas & Mahunka 2001 S04
         |--T. (T.) crassisetosus Willmann 1931 S04
         |    |--T. c. crassisetosus S04
         |    `--T. c. fijiensis Hammer 1971 S04
         |--T. (T.) glaber (Michael 1888) [=Nothrus glaber] S04
         |    |--T. g. glaber S04
         |    `--T. g. javensis Willmann 1931 S04
         |--T. (T.) humeratus Balogh 1997 S04
         |--T. (T.) itatiaiae Balogh 1997 S04
         |--T. (T.) latus (Hammer 1962) S04 [=*Fossonothrus latus BB92]
         |--T. (T.) lisosetosus Iglesias, Palacios-Vargas & Mahunka 2001 S04
         |--T. (T.) magnilamellatus Yamamoto 1996 S04
         |--T. (T.) magnisetosus Iglesias, Palacios-Vargas & Mahunka 2001 S04
         |--T. (T.) novaezealandiae (Hammer 1966) [=Fossonothrus novaezealandiae] S04
         |--T. (T.) pitentzin Iglesias, Palacios-Vargas & Mahunka 2001 S04
         |--T. (T.) saxosus Knülle 1957 [incl. Malaconothrus optatus Berlese 1908 (n. d.)] S04
         |--T. (T.) sculptus Knülle 1957 S04
         |--T. (T.) undulatus Yamamoto, Kuriki & Aoki 1993 S04
         |--T. (T.) wuyanensis Yamamoto, Aoki et al. 1993 S04
         `--T. (T.) yunnanensis Yamamoto & Aoki 1998 S04

Trimalaconothrus (Tyrphonothrus) maior (Berlese 1910) [=Malaconothrus maior; incl. T. intermedius Cooreman 1941, M. novus Sellnick 1921, T. (*Tyrphonothrus) novus, M. sphagnicola Trägardh 1910] S04

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.