Nemesiidae

Lidless banksia trapdoor spider Chenistonia tepperi, copyright William Archer.


Belongs within: Avicularoidea.

The Nemesiidae are a family of trapdoor spiders characterised by a double row of teeth on the paired leg claws.

Characters (from Jocqué & Dippenaar-Schoeman 2007): Medium-sized to very large; cephalic region low; eight eyes present in two rows; palpal claw of female with teeth on promargin; tarsi with scopulae, three tarsal claws present or rarely two, paired claws broad, biserially dentate; four spinnerets present, apical segment of posterior spinnerets digitiform.

<==Nemesiidae [Nemesieae]
    |--Bemmerinae JD-S07
    |--Diplothelopsinae JD-S07
    |--Ixamatus R14 [=Ixalus Koch 1873 (preoc.) R14; Ixamatinae JD-S07]
    |    |--*I. varius [=Ixalus varius, Hapalothele varia] R14
    |    |--I. broomii R14
    |    |--I. distinctus Rainbow 1914 R14
    |    `--I. gregorii R14
    |--Pycnothelinae LP08
    |    |--Chaco LP08
    |    |--Hermachura LP08
    |    |--Prorachias LP08
    |    |--Psalistopoides LP08
    |    |--Pselligmus LP08
    |    |--Pycnothele LP08
    |    |--Rachias LP08
    |    |--Stenoterommata LP08
    |    `--Neostothis Vellard 1925 LP08
    |         `--*N. gigas Vellard 1925 LP08
    |--Nemesia Audouin 1826 [Nemesiinae] JD-S07
    |    |--N. bristowei Decae 2005 F05
    |    |--N. corsica Simon 1914 K55
    |    |--N. dubia K55
    |    |--N. ibiza Decae 2005 F05
    |    |--N. kahmanni Kraus 1955 K55
    |    |--N. randa Decae 2005 F05
    |    |--N. santeugenia Decae 2005 F05
    |    |--N. santeulalia Decae 2005 F05
    |    |--N. seldeni Decae 2005 F05
    |    `--N. valenciae Kraus 1955 K55
    `--Anaminae LP08
         |  i. s.: Acanthogonatus LP08
         |         Longistylus LP08
         |         Hermacha LP08
         `--Anamini MF09
              |--Kwonkan Main 1983 MF09
              |--Swolnpes Main & Framenau 2009 MF09
              |    |--*S. darwini Main & Framenau 2009 MF09
              |    `--S. morganensis Main & Framenau 2009 MF09
              |--Yilgarnia Main 1986 MF09
              |    |--*Y. currycomboides Main 1986 M08
              |    `--Y. linnaei Main 2008 M08
              |--Aname Koch 1873 MF09
              |    |--A. bicolor Rainbow 1914 R14
              |    |--A. mainae MC13
              |    |--A. pallida Koch 1873 R14
              |    `--A. tropica M12
              `--Chenistonia Hogg 1901 [incl. Proshermacha Simon 1908] M12
                   |--*C. maculata Hogg 1901 [=Aname maculata] B12
                   |--C. auropilosa Rainbow & Pulleine 1918 M54
                   |--C. boranup Main 2012 M12
                   |--C. caeruleomontana (Raven 1984) [=Aname caeruleomontana] M12
                   |--C. cuspidata Main 1954 M54
                   |--C. earthwatchorum (Raven 1984) [=Aname earthwatchorum] M12
                   |--C. giraulti Rainbow 1914 M54
                   |--C. hickmani (Raven 1984) [=Aname hickmani] M12
                   |--C. hoggi Rainbow 1914 M54
                   |--C. major Hogg 1901 M54
                   |--C. montana (Raven 1984) [=Aname montana] M12
                   |--*Proshermacha’ subarmata Simon 1908 M12
                   |--C. tepperi Hogg 1902 M54
                   `--C. trevallynia Hickman 1926 M12

Nemesiidae incertae sedis:
  ‘Trechona’ adspersa Bertkau 1880 PCBFF08
  Stanwellia Rainbow & Pulleine 1918 MF09
    `--S. kaituna (Forster 1968) [=Aparua kaituna] NS00
  Cretamygale Selden 2002 S02
    `--*C. chasei Selden 2002 S02
  Scalidognathus Karsch 1892 K92
    `--*S. seticeps Karsch 1892 K92
  Merredinia Main 1983 MF09
  Pseudoteyl Main 1985 MF09
  Teylini MF09
    |--Teyl Main 1975 MF09
    `--Teyloides M12
  Entypesa JD-S07

*Type species of generic name indicated

REFERENCES

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[K92] Karsch, F. 1892. Arachniden von Ceylon und von Minikoy gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift 36 (2): 267–310.

[K55] Kraus, O. 1955. Spinnen von Korsika, Sardinien und Elba (Arach., Araneae). Senckenbergiana Biologica 36: 371–394.

[LP08] Lucas, S. M., V. Passanha, C. R. V. Janini & R. P. Indicatti. 2008. On the genus Neostothis Vellard (Araneae, Nemesiidae). Journal of Arachnology 36 (2): 472–475.

[M54] Main, B. Y. 1954. Spiders and Opiliones. In: Willis, J. H. (ed.) The Archipelago of the Recherche pp. 37–53. Australian Geographical Society: Melbourne.

[M08] Main, B. Y. 2008. A new species of the mygalomorph spider genus Yilgarnia from the Western Australian wheatbelt (Araneae: Nemesiidae). Records of the Western Australian Museum 24 (4): 321–324.

[M12] Main, B. Y. 2012. Description and biogeographic implications of a new species of the Chenistonia maculata group from south-western Western Australia and rediagnosis of Chenistonia (Araneae: Mygalomorphae: Nemesiidae). Records of the Western Australian Museum 27 (2): 176–181.

[MF09] Main, B. Y., & V. W. Framenau. 2009. A new genus of mygalomorph spider from the Great Victoria Desert and neighbouring arid country in south-eastern Western Australia (Araneae: Nemesiidae). Records of the Western Australian Museum 25 (3): 277–285.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[NS00] Nicholls, D. C., P. J. Sirvid, S. D. Pollard & M. Walker. 2000. A list of arachnid primary types held in Canterbury Museum. Records of the Canterbury Museum 14: 37–48.

[PCBFF08] Pedroso, D. R., R. L. Cerqueira Baptista & P. S. Fiúza Ferreira. 2008. Trechona rufa (Araneae, Dipluridae): new status, redescription and neotype designation with notes on the genus. Journal of Arachnology 36 (2): 360–367.

[R14] Rainbow, W. J. 1914. Studies in Australian Araneidae, no. 6, the Terretelariae. Records of the Australian Museum 10 (8): 187–270.

[S02] Selden, P. A. 2002. First British Mesozoic spider, from Cretaceous amber of the Isle of Wight, southern England. Palaeontology 45 (5): 973–983.

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