Cosmochthonius sp., copyright D. E. Walter.

Belongs within: Cosmochthonioidea.

The Cosmochthoniidae are a group of oribatid mites found in dry habitats such as dry soils. They are characterised by having four pairs of large erectile setae arising from transverse scissures on the notogaster. The notogaster is divided by three sutures in Krivolutskiella pubescens, Phyllozetes and Cosmochthonius but only two sutures in Nipponiella simplex (Balogh & Balogh 1992).

Characters (from Norton & Behan-Pelletier 2009): Prodorsum and notogaster usually with reticulate or foveate sculpturing; rostrum distinctly fenestrate. Base of bothridial seta with conspicuous sharp proximal d bend. Setae of rows cf occupying anterior half of notogaster; notogaster with threee transeverse scissures, such that rows c and d on separate plates, setal rows c and d not enlarged; rows e and f much longer than others, inserted on intercalary sclerites in two type S scissures. Opisthonotal gland absent. Genital plate not divided; aggenital plates small, not fused to coxisternum; adanal plates fused posteriorly to form a U shape. Legs with five free segments past coxisternum (femora undivided); pretarsi of legs bi- or tridactylous.

Cosmochthoniidae [Cosmochthoniides]
    |--Krivolutskiella Gordeeva 1980 S04
    |    `--*K. pubescens Gordeeva 1980 S04
    |--Nipponiella Gordeeva 1980 S04
    |    `--*N. simplex (Aoki 1966) [=Trichthonius simplex] S04
    |--Phyllozetes Gordeeva 1978 S04
    |    |--*P. emmae (Berlese 1910) S04 (see below for synonymy)
    |    |--P. alatus Mahunka 1982 S04
    |    |--P. hypoquercus McDaniel & Bolen 1980 S04
    |    |--P. latifolius Gordeeva 1980 S04
    |    |--P. longifolius Balakrishnan 1986 S04
    |    |--P. nilamburicus Balakrishnan 1986 S04
    |    |--P. osithchnjukovi Gordeeva 1980 S04
    |    `--P. tauricus Gordeeva 1978 S04
    `--Cosmochthonius Berlese 1910 S04
         |--C. (Cosmochthonius) S04
         |    |--*C. (C.) lanatus (Michael 1885) [=Hypochthonius lanatus] S04
         |    |    |--C. l. lanatus [incl. C. arctosus Pankov 2002, C. wallworki Lee 1982] S04
         |    |    |--C. l. diversiseta Sarkar & Subías 1982 S04
         |    |    `--C. l. foveolatus Beck 1962 S04
         |    |--C. (C.) asiaticus Gordeeva 1980 S04
         |    |--C. (C.) assamensis Talukdar & Chakrabarti 1984 S04
         |    |--C. (C.) australicus Womersley 1945 [=C. plumatus australicus] S04
         |    |--C. (C.) bengalensis Chakrabarti, Bhaduri & Raychaudhuri 1972 S04
         |    |--C. (C.) bhutanensis Chakrabarti & Wilson 1981 S04
         |    |--C. (C.) concavus Aoki 1994 S04
         |    |--C. (C.) desaussurei Mahunka 1982 S04
         |    |--C. (C.) domesticus Grandjean 1947 (n. d.) S04
         |    |--C. (C.) foliatus Subías 1982 S04
         |    |--C. (C.) imperfectus Aoki 2000 S04
         |    |--C. (C.) macrosetosus Ayyildiz & Luxton 1990 S04
         |    |--C. (C.) maroccanus Gil-Martín, Subías & Arillo 1992 S04
         |    |--C. (C.) minifoveolatus Gil-Martín, Subías Candelas 1991 S04
         |    |--C. (C.) monegrensis Pérez-Íñigo 1991 S04
         |    |--C. (C.) nayoroensis Fujikawa 1980 S04
         |    |--C. (C.) novus Sergienko 1991 S04
         |    |--C. (C.) pallidus Lombardini 1961 (n. d.) S04
         |    |--C. (C.) perezinigoi Morell 1988 S04
         |    |--C. (C.) plumatus Berlese 1910 S04
         |    |    |--C. p. plumatus S04
         |    |    `--C. p. suramericanus Hammer 1958 S04
         |    |--C. (C.) ponticus Gordeeva 1980 S04
         |    |--C. (C.) reticulatus Grandjean 1947 S04, NB-P09, S04
         |    |--C. (C.) semiareolatus Hammer 1966 S04
         |    |--C. (C.) semifoveolatus Subías 1982 S04
         |    |--C. (C.) signatus Pérez-Íñigo 1990 S04
         |    |--C. (C.) spinosus Gil-Martín, Subías & Candelas 1991 S04
         |    |--C. (C.) sublanatus Mahunka 1977 S04
         |    |--C. (C.) taurus Niemi, Gordeeva & Ayyildiz 2002 S04
         |    |--C. (C.) tenuisetus Gordeeva 1980 S04
         |    |--C. (C.) trivialis Sergienko 1991 S04
         |    `--C. (C.) ugamaensis Gordeeva 1980 S04
         `--C. (Nanochthonius Subías & Gil-Martín 1995) (see below for synonymy) S04
              |--C. (*N.) ruizi Kahwash, Subías & Ruiz 1989 S04 [=C. (*Microchthonius) ruizi S04, BB92]
              `--C. (N.) juvenalis Kamill 1986 S04

Cosmochthonius (Nanochthonius Subías & Gil-Martín 1995) [=Microchthonius Kahwash, Subías & Ruiz 1989 non Hadzi 1933] S04

*Phyllozetes emmae (Berlese 1910) S04 [=Cosmochthonius emmae S04, Ovochthonius emmae CH98, Phylozetes (l. c.) emmae H98; incl. Trichthonius heterotrichus Sanyal & Bhaduri 1983 S04]

*Type species of generic name indicated


[BB92] Balogh, J. & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 13 January 2019.

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