Atropacarus sp., copyright Scott Justis.

Belongs within: Phthiracaridae.

Atropacarus is a cosmopolitan genus of phthiracarid mites with three or four pairs of closely-placed setae on the inner margin of the ano-adanal plates. The type subgenus Atropacarus, with four pairs of such setae, is sometimes treated separately from the remaining subgenera with three pairs, which are then placed as the genus Hoplophorella.

<==Atropacarus Ewing 1917 [incl. Helvetacarus Mahunka 1993] S04
    |  i. s.: A. bichei Niedbala 1986 S04
    |         A. ciliosus Niedbala 2002 S04
    |         A. clavatus Aoki 1980 [=A. sticulus clavatus] S04
    |         A. clavigerus (Berlese 1904) [=Hoploderma clavigerum; incl. Phthiracarus lutulentus Sellnick 1920] S04
    |         A. controversus Niedbala 2000 S04
    |         A. csiszarae (Balogh & Mahunka 1979) [=Steganacarus csiszarae] S04
    |         A. echinodiscus (Mahunka 1982) [=Steganacarus echinodiscus] S04
    |         A. genavensis (Mahunka 1993) [=Helvetacarus genavensis] S04
    |         A. inconditus Mahunka 1991 S04
    |         A. macrosculpturatus (Mahunka & Mahunka-Papp 1999) [=Steganacarus macrosculpturatus] S04
    |         A. mirabilis (Mahunka 1979) [=Steganacarus mirabilis] S04
    |         A. multisetosus Niedbala 2002 S04
    |         A. murciensis Niedbala 1986 S04
    |         A. paraclavatus Niedbala 2002 S04
    |         A. phyllophorus (Berlese 1904) (see below for synonymy) S04
    |         A. platakisi (Mahunka 1979) (see below for synonymy) S04
    |         A. punctulatus Moraza 1985 S04
    |         A. serratus (Feider & Suciu 1957) [=Steganacarus serratus] S04
    |         A. terrapene (Jacot 1937) [=Steganacarus terrapene] S04
    |         A. wandae (Niedbala 1981) [=Steganacarus wandae; incl. S. navarrensis Moraza 1985] S04
    |--A. (Atropacarus) S04
    |    |--*A. (A.) striculus (Koch 1835) [=Hoplophora stricula] S04
    |    |    |--A. s. striculus (see below for synonymy) S04
    |    |    `--A. s. insularis (Weigmann 1976) [=Steganacarus insularis] S04
    |    |--A. (A.) absimilis (Niedbala 1982) [=Steganacarus (A.) absimilis] S04
    |    |--A. (A.) decipiens (Niedbala 1984) [=Steganacarus (A.) decipiens] S04
    |    |--A. (A.) folious Niedbała 2003 N07
    |    |--A. (A.) griseus (Niedbala 1984) [=Steganacarus (A.) griseus] S04
    |    |--A. (A.) immundus (Niedbala 1983) [=Steganacarus (A.) immundus] S04
    |    |--A. (A.) inculpatus (Niedbala 1984) [=Steganacarus (A.) inculpatus] S04
    |    |--A. (A.) maculosus (Niedbala 1983) [=Steganacarus (A.) maculosus] S04
    |    |--A. (A.) obesus (Niedbala 1983) [=Steganacarus (A.) obesus] S04
    |    |--A. (A.) ochraceus (Niedbala 1983) [=Steganacarus (A.) ochraceus] S04
    |    |--A. (A.) parvulus (Niedbala 1983) [=Steganacarus (A.) parvulus] S04
    |    |--A. (A.) perversus (Niedbala 1983) [=Steganacarus (A.) perversus] S04
    |    `--A. (A.) substrictus (Niedbala 1983) [=Steganacarus (A.) substrictus] S04
    |--H. (Hoplophorella Berlese 1923) N07, S04
    |    |--A. (*H.) cucullatus (Ewing 1909) BB92, S04 (see below for synonymy)
    |    |    |--A. c. cucullatus (see below for synonymy) S04
    |    |    `--‘Hoplophorella’ c. curassensis Willmann 1936 S04
    |    |--‘Hoplophorella’ (H.) angolensis Mahunka 1984 S04
    |    |--‘Hoplophorella’ (H.) berninii (Niedbala 1991) [=Protophthiracarus berninii] S04
    |    |--‘Hoplophorella’ (H.) brasiliensis (Niedbala 1988) [=Austrophthiracarus brasiliensis] S04
    |    |--‘Hoplophorella’ (H.) brevipilis (Balogh 1958) [=Steganacarus brevipilis] S04
    |    |--‘Hoplophorella’ (H.) collaris (Balogh 1958) [=Steganacarus collaris] S04
    |    |--‘Hoplophorella’ (H.) craterifer (Hammer 1971) [=Steganacarus craterifer] S04
    |    |--A. (H.) diaphoros Niedbala 2000 S04
    |    |--‘Hoplophorella’ (H.) digna Niedbala 1983 S04
    |    |--A. (H.) dissimilis Niedbala 1998 S04
    |    |--‘Hoplophorella’ (H.) dubia (Niedbala 1992) [=Hoplophthiracarus dubius] S04
    |    |--‘Hoplophorella’ (H.) ensifera Mahunka 1985 S04
    |    |--‘Hoplophorella’ (H.) ephippigera (Balogh & Mahunka 1977) [=Steganacarus ephippiger] S04
    |    |--‘Hoplophorella’ (H.) evergladensis (Niedbala 2001) [=Protophthiracarus evergladensis] S04
    |    |--‘Hoplophorella’ (H.) fonseciai (Pérez-Íñigo & Baggio 1980) [=Steganacarus fonseciai] S04
    |    |--A. (H.) frondeus Niedbała 2003 N07
    |    |--‘Hoplophorella’ (H.) grandjeani Niedbala 1981 S04
    |    |--A. (H.) hamatus (Ewing 1909) N07 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) heterotricha Mahunka 1984 S04
    |    |--‘Hoplophorella’ (H.) insolens Neidbala 1988 S04
    |    |--‘Hoplophorella’ (H.) kulczynskii Niedbala 1981 S04
    |    |--‘Hoplophorella’ (H.) lanceoseta (Balogh & Mahunka 1981) (see below for synonymy) S04
    |    |--‘Hoplophorella’ (H.) lanceosetoides Mahunka 1985 S04
    |    |--‘Hoplophorella’ (H.) lemuria Mahunka 1993 S04
    |    |--‘Hoplophorella’ (H.) manipurensis Misra, Bhaduri & Raychaudhuri 1982 S04
    |    |--‘Hoplophorella’ (H.) minisetosa Mahunka 1978 S04
    |    |--‘Hoplophorella’ (H.) neonominata Subías 2004 (see below for synonymy) S04
    |    |--A. (H.) nigeriensis Badejo, Woas & Beck 2001 S04
    |    |--‘Hoplophorella’ (H.) oblonga Niedbala 1983 S04
    |    |--A. (H.) pallens (Niedbala 1983) N91 [=Hoplophorella pallens S04]
    |    |--‘Hoplophorella’ (H.) paraensis (Pérez-Íñigo & Baggio 1996) [=Protophthiracarus paraensis] S04
    |    |--A. (H.) pergratus Niedbala 1998 [=Hoplophorella pergrata] S04
    |    |--‘Hoplophorella’ (H.) perisi Subías & Sarkar 1984 S04
    |    |--‘Hoplophorella’ (H.) pervicax (Niedbala 1984) [=Hoplophthiracarus pervicax] S04
    |    |--‘Hoplophorella’ (H.) praeoccupata Subías 2004 (see below for synonymy) S04
    |    |--‘Hoplophorella’ (H.) prominens (Balogh 1958) [=Steganacarus prominens] S04
    |    |--‘Hoplophorella’ (H.) pustulata Mahunka 1993 S04
    |    |--A. (H.) rafalskii (Niedbala 1997) N91, S04 [=Notophthiracarus rafalskii S04]
    |    |--‘Hoplophorella’ (H.) rangiroaensis Hammer 1972 S04
    |    |--‘Hoplophorella’ (H.) remigera (Berlese 1923) [=Phthiracarus (Trachyhoplophora) remiger] S04
    |    |--‘Hoplophorella’ (H.) repetita Subías 2004 (see below for synonymy) S04
    |    |--‘Hoplophorella’ (H.) sabahna Mahunka 1991 S04
    |    |--‘Hoplophorella’ (H.) sculptilis (Niedbala 1988) (see below for synonymy) S04
    |    |--A. (H.) singularis (Sellnick 1959) N91, S04 (see below for synonymy)
    |    |--A. (H.) stilifer (Hammer 1961) N91, S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) subciliata Mahunka 1983 S04
    |    |--‘Hoplophorella’ (H.) subita Niedbala 1983 S04
    |    |--‘Hoplophorella’ (H.) subsellata Balogh & Mahunka 1977 (see below for synonymy) S04
    |    |--‘Hoplophorella’ (H.) tripartita (Niedbala & Schatz 1996) [=Protophthiracarus tripartitus] S04
    |    |--‘Hoplophorella’ (H.) unqus (Niedbala 1991) [=Notophthiracarus unqus] S04
    |    |--‘Hoplophorella’ (H.) varia (Niedbala & Schatz 1996) [=Protophthiracarus varius] S04
    |    |--‘Hoplophorella’ (H.) varians Jacot 1933 S04
    |    |--‘Hoplophorella’ (H.) venusta Niedbala 1983 S04
    |    `--A. (H.) vitrinus (Berlese 1913) N01, S04 (see below for synonymy)
    |--‘Hoplophorella’ (Kakophthiracarus Mahunka 1992) S04
    |    |--‘Hoplophorella’ (K.) tuberculosissima (Mahunka 1978) (see below for synonymy) S04
    |    |--‘Hoplophorella’ (K.) balazsi Mahunka 1983 S04
    |    |--‘Hoplophorella’ (K.) cavernosa Mahunka 1987 S04
    |    |--‘Hoplophorella’ (K.) enigmatica (Mahunka 1996) [=Kakophthiracarus enigmaticus] S04
    |    |--‘Hoplophorella’ (K.) multirugosa (Mahunka 1978) (see below for synonymy) S04
    |    |--‘Hoplophorella’ (K.) multituberculata (Balogh & Mahunka 1966) [=Steganacarus multituberculatus] S04
    |    `--‘Hoplophorella’ (K.) mwali (Mahunka 1994) [=Kakophthiracarus mwali] S04
    `--‘Hoplophorella’ (Steganacarellus Mahunka 1986) S04
         |--‘Hoplophorella’ (*S.) reducta (Mahunka 1986) [=*Steganacarellus reductus] S04
         |--‘Hoplophorella’ (S.) heterosetosa (Niedbala 1988) [=Notophthiracarus heterosetosus] S04
         |--‘Hoplophorella’ (S.) ‘heterotricha’ (Niedbala 2000) (see below for synonymy) S04
         |--‘Hoplophorella’ (S.) paraparvula (Niedbala 2000) [=Notophthiracarus paraparvulus] S04
         |--‘Hoplophorella’ (S.) parvula (Niedbala 1998) [=Notophthiracarus parvulus] S04
         `--‘Hoplophorella’ (S.) perlucunda (Niedbala 2000) [=Notophthiracarus perlucundus] S04

Atropacarus phyllophorus (Berlese 1904) [=Hoploderma phyllophorum; incl. Steganacarus ropalus Feider & Suciu 1957] S04

Atropacarus platakisi (Mahunka 1979) [=Steganacarus platakisi; incl. A. papillatus Gil-Martín, Subías & Arillo 1992] S04

*Atropacarus (Atropacarus) striculus striculus (Koch 1835) [incl. Steganacarus diaphanus Jacot 1930, S. senex Aoki 1958, S. trichosus Mahunka 1982] S04

Atropacarus (*Hoplophorella) cucullatus (Ewing 1909) BB92, S04 [=Hoploderma cucullatum S04, Hoplophora cucullatum BB92, *Hoplophorella cucullata S04]

Atropacarus (*Hoplophorella) cucullatus cucullatus (Ewing 1909) [incl. Hoplophorella cucullata cuculloides Jacot 1933, Hoploderma licnophorum Berlese 1913] S04

Atropacarus (Hoplophorella) hamatus (Ewing 1909) N07 [=Hoploderma hamatum S04, Hoplophorella hamata S04; incl. Hoplophorella cochlearia Pérez-Íñigo & Baggio 1993 S04, Hoplophorella cuneiseta Mahunka 1988 S04, Hoplophorella cucullata floridae Jacot 1933 S04, Atropacarus (Hoplophorella) floridae N91, Hoplophorella glauca Hammer 1972 S04, A. (H.) glaucus N91, Hoplophorella ligulifera Mahunka 1987 S04, Phthiracarus (H.) cucullatus obsoletior Berlese 1923 S04, Hoplophorella schauenbergi Mahunka 1978 S04, Atropacarus (H.) schauenbergi N91, Hoplophorella spatulata Parry 1980 S04]

Atropacarus (Hoplophorella) singularis (Sellnick 1959) N91, S04 [=Hoplophorella singularis S59; incl. H. queenslandicus Balogh & Mahunka 1978 S04, H. regalis Mahunka 1978 S04, H. sundarbanensis Sanyal & Bhaduri 1982 S04]

Atropacarus (Hoplophorella) stilifer (Hammer 1961) N91, S04 [=Steganacarus stilifer S04, Hoplophorella (H.) stilifera S04; incl. Hoplophorella benoiti Mahunka 1984 S04]

Atropacarus (Hoplophorella) vitrinus (Berlese 1913) N01, S04 [=Hoploderma vitrinum S04, Hoplophorella vitrina S04; incl. Hoplophorella africana Wallwork 1967 S04, Steganacarus andrei Balogh 1958 S04, Hoplophorella lienhardi Mahunka 1987 S04, H. raychaudhurii Subías & Sarkar 1984 S04, H. scapellata Aoki 1965 S04, Atropacarus (Hoplophorella) scapellatus N91]

‘Hoplophorella’ (Hoplophorella) lanceoseta (Balogh & Mahunka 1981) [=Steganacarus lanceosetus; incl. H. neglecta Niedbala 1984] S04

‘Hoplophorella’ (Hoplophorella) neonominata Subías 2004 [=Protophthiracarus cavernosus Niedbala 2002 non H. cavernosa Mahunka 1987] S04

‘Hoplophorella’ (Hoplophorella) praeoccupata Subías 2004 [=Atropacarus (H.) tuberosus Niedbala & Schatz 1996 non H. tuberosa Mahunka 1988] S04

‘Hoplophorella’ (Hoplophorella) repetita Subías 2004 [=Protophthiracarus reductus Niedbala 1991 nec Steganacarellus reductus Mahunka 1986 nec Hoplophorella reducta Mahunka 1988] S04

‘Hoplophorella’ (Hoplophorella) sculptilis (Niedbala 1988) [=Hoplophthiracarus sculptilis; incl. Hoplophthiracarus magnus Mahunka 1988] S04

‘Hoplophorella’ (Hoplophorella) subsellata Balogh & Mahunka 1977 [incl. Steganacarus galeatus Balogh & Mahunka 1978] S04

‘Hoplophorella’ (Kakophthiracarus) multirugosa (Mahunka 1978) [=Steganacarus multirugosus; incl. S. cornutus Mahunka 1978] S04

‘Hoplophorella’ (Kakophthiracarus) tuberculosissima (Mahunka 1978) [=Steganacarus tuberculosissimus; incl. H. horrida Mahunka 1984, H. meszarosi Mahunka 1988, H. (*K.) tuberosa Mahunka 1988, H. verrucosa Mahunka 1987] S04

‘Hoplophorella’ (Steganacarellus) ‘heterotricha’ (Niedbala 2000) [=Arphthicarus heterotrichus non H. heterotricha Mahunka 1984] S04

*Type species of generic name indicated


[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[N91] Niedbala, W. 1991. Preliminary hypotheses on the biogeography of the Phthiracaroidea (Acari, Oribatida). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 219–231. SPB Academic Publishing: The Hague.

[N01] Niedbała, W. 2001. Study of the diversity of Ptyctima (Acari: Oribatida) and quest for centres of its origin: the fauna of the Oriental and Australian regions. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 143–148. CSIRO Publishing: Melbourne.

[N07] Niedbała, W. 2007. Ptyctimous mites (Acari, Oribatida) of Costa Rica. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 39–42. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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