Haplochthonius simplex, copyright Mitsunobu Shibata.

Belongs within: Enarthronota.

The Haplochthoniidae are a group of somewhat paedomorphic oribatid mites found in dry habitats such as building debris or dry soils.

Characters (from Norton & Behan-Pelletier 2009): Body legnth little more than twice maximum width. Prodorsum stegasime, without naso; bothridial setae clavate, with conspicuous sharp proximal d bend. Legs with five free segments past coxisternum, femora undivided; legs II and III almost adjacent. Pretarsi with empodium developed as claw. Notogaster with three transverse type E scissures; four pairs of setae (c row only) on anteriormost plate; with cupule immediately lateral to each mid-dorsal seta (c1, d1, e1, f1) in addition to normal complement; opisthonotal gland absent. Genital plates twice size of anal plates; aggenital setae absent; anal plates with four pairs of setae; peranal plate and seta absent.

    |--Amnemochthonius Grandjean 1949 S04
    |    `--*A. taeniophorus Grandjean 1948 S04
    `--Haplochthonius Willmann 1930 [incl. Tetrochthonius Hammer 1958] S04
         |--H. (Haplochthonius) S04
         |    |--*H. (H.) simplex (Willmann 1930) [=Cosmochthonius (*H.) simplex] S04
         |    |--H. (H.) antarcticus Sanyal, Basak & Barman 2002 S04
         |    |--H. (H.) chamela Mahunka & Mejía-Recamier 1998 S04
         |    |--H. (H.) clavatus (Hammer 1958) S04 [=*Tetrochthonius clavatus SS82]
         |    |--H. (H.) crassisetosus Gil-Martín, Subías & Arillo 1992 S04
         |    |--H. (H.) graecus Mahunka 1992 S04
         |    |--H. (H.) intermedius Chakrabarti, Bhaduri & Raychaudhuri 1977 S04
         |    |--H. (H.) longisetus Sanyal, Basak & Barman 2002 S04
         |    |--H. (H.) maitri Sanyal, Basak & Barman 2002 S04
         |    |--H. (H.) muscicola Fujikawa 1996 S04
         |    |--H. (H.) sanctaeluciae Bernini 1973 S04
         |    `--H. (H.) variabilis Wallwork 1972 S04
         `--H. (Senilochthonius Mahunka 1992) S04
              `--H. (*S.) baobab (Mahunka 1992) [=*Senilochthonius baobab] S04

*Type species of generic name indicated


Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[SS82] Sarkar, S., & L. S. Subías. 1982. Some new macropylines oribates (Acarida) from India (Hypochthoniidae, Cosmochthonoidea and Epilohmanniidae). Eos 58: 311–318.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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