Aeroppia

Dorsal (left) and ventral views of Aeroppia peruensis, from Balogh & Balogh (1992 vol. 2).

Belongs within: Oppiidae.

Aeroppia is a genus of oppiid oribatid mites found in the Americas.

Characters (from Balogh & Balogh 1992): Prodorsum with costulae, lamellar and translamellar lines absent; interlamellar setae well developed; sensillus capitate. Notogaster with twelve pairs of setae; setae p1 large, more or less dilated; crista absent. Genital plates with five pairs of setae.

<==Aeroppia Hammer 1961 S04
    |--*A. peruensis Hammer 1961 S04
    |--A. abdita Pérez-Íñigo & Baggio 1997 S04
    |--A. adjacens Mahunka 1985 S04
    |--A. asymmetrica Mahunka 1985 S04
    |--A. clavatum Higgins 1966 S04
    |--A. floridana (Banks 1896) [=Belba floridana; incl. Oribata consimilis Banks 1910] S04
    |--A. hammerae Mahunka 1985 S04
    |--A. insularis Higgins 1966 S04
    |--A. magnipilosa (Ewing 1909) [=Damaeus magnipilosus; incl. A. columbiana Hammer 1961] S04
    |--A. mariehammerae Subías et al. 2004 S04
    |--A. nasalis Mahunka 1984 S04
    |--A. sculpturata Mahunka 1985 S04
    `--A. vacua (Berlese 1888) [=Belba concolor vacua] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Neoamerioppia

Dorsal (left) and ventral views of Neoamerioppia decemsetosa, from Balogh & Balogh (1992).

Belongs within: Oppiidae.

Neoamerioppia is a pantropical and subtropical genus of oppiid oribatid mites in which the interlamellar setae are absent.

<==Neoamerioppia Subías 1989 S04
    |--N. (Neoamerioppia) S04
    |    |--*N. (N.) decemsetosa (Hammer 1973) [=Amerioppia decemsetosa] S04
    |    |--N. (N.) aelleni (Mahunka 1982) [=Amerioppia aelleni] S04
    |    |--N. (N.) africana (Kok 1967) [=Amerioppia africana] S04
    |    |--N. (N.) asiatica (Hammer 1977) [=Amerioppia asiatica] S04
    |    |--N. (N.) chavinensis (Hammer 1961) [=Amerioppia chavinensis] S04
    |    |--N. (N.) chilensis (Hammer 1962) [=Amerioppia chilensis] S04
    |    |--N. (N.) cocuyana (Balogh 1984) [=Amerioppia cocuyana] S04
    |    |--N. (N.) deficiens (Balogh 1959) [=Oppia deficiens] S04
    |    |--N. (N.) extrusa (Mahunka 1983) [=Amerioppia extrusa] S04
    |    |--N. (N.) flagellata (Hammer 1975) [=Amerioppia flagellata] S04
    |    |--N. (N.) flagelliapex Ohkubo & Aoki 1995 S04
    |    |--N. (N.) foveolata (Mahunka 1984) [=Amerioppia foveolata] S04
    |    |--N. (N.) hamidi (Al-Assiuty & El-Deeb 1983) [=Amerioppia hamidi] S04
    |    |--N. (N.) hexapilis (Hammer 1961) [=Amerioppia hexapilis] S04
    |    |--N. (N.) interrogata (Mahunka 1976) [=Amerioppia interrogata] S04
    |    |--N. (N.) lanceolata (Hammer 1958) [=Oppia lanceolata] S04
    |    |--N. (N.) longiclava (Hammer 1962) [=Amerioppia longiclava] S04
    |    |    |--N. l. longiclava S04
    |    |    `--N. l. microseta (Balogh & Balogh 1986) [=Amerioppia longiclava microseta] S04
    |    |--N. (N.) longicoma (Hammer 1958) [=Oppia longicoma] S04
    |    |--N. (N.) minima (Hammer 1961) [=Amerioppia minima] S04
    |    |--N. (N.) nagyi (Mahunka 1969) [=Oppia nagyi] S04
    |    |--N. (N.) notata (Hammer 1958) [=Oppia notata] S04
    |    |--N. (N.) papuana (Balogh & Balogh 1986) [=Amerioppia papuana] S04
    |    |--N. (N.) paripilis (Hammer 1961) [=Amerioppia paripilis] S04
    |    |--N. (N.) pectigera (Hammer 1961) [=Amerioppia pectigera] S04
    |    |--N. (N.) phoretica (Franklin & Woas 1992) [=Oppia phoretica] S04
    |    |--N. (N.) polygonata (Mahunka 1982) [=Amerioppia polygonata] S04
    |    |--N. (N.) rotunda (Hammer 1958) [=Oppia rotunda] S04
    |    |--N. (N.) similis (Covarrubias 1967) [=Amerioppia similis] S04
    |    |--N. (N.) sturmi (Balogh 1984) [=Amerioppia sturmi] S04
    |    |--N. (N.) trichosa (Hammer 1958) [=Oppia trichosa] S04
    |    |--N. (N.) trichosoides (Hammer 1961) [=Amerioppia trichosoides] S04
    |    |--N. (N.) ventrosquamosa (Hammer 1979) [=Amerioppia ventrosquamosa] S04
    |    |--N. (N.) vicina (Hammer 1971) [=Amerioppia vicina] S04
    |    |--N. (N.) vietnamica (Mahunka 1988) [=Amerioppia vietnamica] S04
    |    `--N. (N.) woolleyi (Hammer 1968) [=Amerioppia woolleyi] S04
    `--N. (Amerigloboppia Subías 1989) S04
         |--N. (*A.) espeletiarum (Balogh 1984) [=Amerioppia espeletiarum] S04
         |--N. (A.) badensis (Woas 1986) [=Oppia badensis] S04
         |--N. (A.) centraliamericana (Mahunka 1983) [=Globoppia centraliamericana] S04
         |--N. (A.) costulifera Mahunka 1997 S04
         |--N. (A.) extrema (Mahunka 1985) [=Amerioppia extrema] S04
         |--N. (A.) salvadoriensis (Woas 1986) [=Oppia salvadoriensis] S04
         `--N. (A.) senecionis (Balogh 1984) [=Amerioppia senecionis] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Taiwanoppia

Dorsal (left) and ventral views of Taiwanoppia hungarorum, from Balogh & Balogh (1992 vol. 2).

Belongs within: Oppiidae.

Taiwanoppia is a pantropical genus of smooth-bodied oppioid oribatid mites.

Characters (from Balogh & Balogh 1992, as Vietoppia): Prodorsum with costulae, lamellar and translamellar lines absent; interlamellar setae well developed; sensillus capitate. Body smooth; notogaster with nine pairs of setae; without protruding humeral processes; crista absent. Apodemata IV present. Genital plates with five pairs of setae.

<==Taiwanoppia Tseng 1982 S04
    |--T. (Taiwanoppia) [incl. Vietoppia Mahunka 1988] S04
    |    |--*T. (T.) subtropica Tseng 1982 S04
    |    |--T. (T.) fujianensis (Wang 1993) [=Vietoppia fujianensis] S04
    |    |--T. (T.) hungarorum (Mahunka 1988) S04 [=*Vietoppia hungarorum BB92]
    |    `--T. (T.) papillaris Tseng 1982 S04
    `--T. (Paragloboppia Subías 1989) S04
         |--T. (*P.) diversiseta (Mahunka 1985) S04 [=Oppia diversiseta S04, Vietoppia (*P.) diversiseta BB92]
         |--T. (P.) calva (Warburton 1912) [=Notaspis clavipectinata calva] S04
         |--T. (P.) mercedesae (Arillo & Subías 1997) [=Vietoppia (P.) mercedesae] S04
         |--T. (P.) pacifica (Ohkubo & Aoki 1995) [=Vietoppia (P.) pacifica] S04
         |--T. (P.) senegalensis (Mahunka 1975) [=Oppia senegalensis] S04
         `--T. (P.) trichotos (Balogh & Mahunka 1977) [=Oppia trichotos] S04

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Oppia

Oppia nitens, from here.

Belongs within: Oppioidea.

Oppia is a genus of oppioid oribatid mites found in most parts of the world except Australia (Subías 2004). They are fungal feeders with masses of fungal spores having been found in the gut of O. nitens and O. neerlandica (Norton & Behan-Pelletier 2009).

Characters (from Balogh & Balogh 1992): Prodorsum with costulae, lamellar and translamellar lines absent; interlamellar setae present; sensillus elongately fusiform or lanceolate. Notogaster without protruding humeral processes; crista absent. Genital plate with five pairs of setae.

Oppia Koch 1836 [incl. Dameosoma Berlese 1892] S04
    |--*O. nitens Koch 1836 S04
    |    |--O. n. nitens (see below for synonymy) S04
    |    `--O. n. brachytrichinus Dalenius 1958 S04
    |--O. alces (Jacot 1934) SN98
    |--O. capense (Paoli 1908) [=Dameosoma capense] S04
    |--O. cephalota (Hall 1911) (n. d.) [=Damaeus cephalotus] S04
    |--O. coloradensis Woolley 1969 S04
    |--O. concolor Koch 1839 (n. d.) S04
    |--O. denticulata (Canestrini & Canestrini 1882) S04 (see below for synonymy)
    |--O. elegans (Trägårdh 1931) (n. d.) [=Dameosoma elegans] S04
    |--O. elongata Jacot 1938 (n. d.) [incl. O. parviaures Jacot 1939] S04
    |--O. ewingi (Berlese 1916) S04 [=Damoeosoma denticulatum var. ewingi S04]
    |--O. falcata N77
    |--O. flagellifera Wang 1993 S04
    |--O. huangshanensis Wen & Chen 1991 S04
    |--O. hungarica Karpelles 1893 (n. d.) S04
    |--O. incisirostra Woas 1986 S04
    |--O. leleupi Balogh 1958 (n. d.) S04
    |--O. neerlandica NB-P09 [=Dameosoma neerlandicum CH98]
    |--O. neonominata Subías 2004 [=Oppia concolor tridentata Pérez-Íñigo 1976 non O. tridentata Forsslund 1942] S04
    |--O. obsoleta N77
    |--O. orientalis Sanyal & Bhaduri 1989 S04
    |--O. ornata (Oudemans 1900) S62
    |--O. quadrisetosa (Kramer 1898) (n. d.) [=Notaspis quadrisetosa] S04
    |--O. samadi Kardar 1976 S04
    |--O. seminuda Scull, Jeleva & Cruz 1984 S04
    |--O. speciosa Golosova 1981 S04
    |--‘Dameosoma’ splendens S22
    |--O. unicarinata N77
    `--O. vitrea (Tseng 1982) [=Brachioppia vitrea] S04

Oppia denticulata (Canestrini & Canestrini 1882) S04 [=Belba denticulata S04, *Dameosoma denticulatum CH98; incl. O. cyclosoma Mihelčič 1955 S04, O. grandis Mihelčič 1955 S04, O. willmanni Oudemans 1937 S04]

*Oppia nitens nitens Koch 1836 [incl. Belba minuta Banks 1895, Dameosoma myrmecophila Sellnick 1928, Oribata perolata Banks 1909] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[N77] Niedbała, W. 1977. Badania biocenotyczne nad mechowcami (Acari, Oribatei) Masywu Ślęży. Badania Fizjograficzne nad Polską Zachodnią, Seria C—Zoologia 30: 47–73.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S62] Seyd, E. L. 1962. The moss mites of Kinder Scout, Derbyshire (Acari: Oribatei). Journal of the Linnean Society of London, Zoology 44 (300): 585–591.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[SN98] Swift, S. F., & R. A. Norton. 1998. Preliminary report on oribatid mite (Acari: Oribatida) diversity in the Hawaiian Islands. Bishop Museum Occasional Papers 57: 1–44.

Last updated: 22 December 2018.

Machuellidae

Dorsal (left) and ventral views of Machuella sp., from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

The Machuellidae are a group of oppioid oribatid mites in which the epimeral setae form a ventral basket covered by cerotegument.

Characters (from Norton & Behan-Pelletier 2009): Length 150–250 µm. Coxisternal setal formula 2-2-4-5 or up to seventeen pairs of coxisternal setae, coxisternal setae directed medially toward ventrosejugal region, forming a "basket" within a thick layer of cerotegument; anterior margin of epimere I with medial tooth. Distance between genital and anal plates less than length of genital plate.

<==Machuellidae [Machuellinae] S04
    |--Gredosella Gil-Martín, Arillo & Subías 2000 S04
    |    |--*G. fraternalis Gil-Martín, Arillo & Subías 2000 S04
    |    `--G. capitata (Kulijev 1967) [=Machuella capitata] S04
    `--Machuella Hammer 1961 S04
         |--*M. ventrisetosa Hammer 1961 S04
         |    |--M. v. ventrisetosa [incl. M. hellenica Mahunka 1982, M. hippy Niemi & Gordeeva 1991] S04
         |    |--M. v. bilineata Weigmann 1976 S04
         |    |--M. v. plicata Hammer 1979 S04
         |    |--M. v. pyriformis Hammer 1968 S04
         |    |--M. v. robusta Hammer 1971 S04
         |    `--M. v. zehntneri Mahunka 1977 S04
         |--M. draconis Hammer 1961 [incl. M. africana Mahunka 1978] S04
         `--M. lineata Hammer 1973 S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Sternoppia

Dorsal (left) and ventral views of Sternoppia brasiliensis, from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

Sternoppia is a genus of oppioid oribatid mites characterised by large plumose coxisternal setae together with two overlapping plates covering the coxisternum (Norton & Behan-Pelletier 2009).

<==Sternoppia Balogh & Mahunka 1968 [incl. Synoppia Balogh & Mahunka 1969; Sternoppiidae] S04
    |--*S. mirabilis Balogh & Mahunka 1968 S04
    |--S. boliviana Balogh & Mahunka 1969 S04
    |--S. brasiliensis Franklin & Woas 1992 S04
    |--S. incisa Balogh & Mahunka 1977 S04
    |--S. minor Balogh & Mahunka 1980 S04
    |--S. quadriseta (Balogh & Mahunka 1969) [=Synoppia quadriseta] S04
    |--S. reticulata Balogh & Mahunka 1969 S04
    |--S. sphaerodendron Balogh & Mahunka 1979 S04
    |--S. striata Mahunka 1983 S04
    `--S. vicina Balogh & Mahunka 1980 S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Quadroppiidae

Quadroppia quadricarinata, from here.

Belongs within: Circumdehiscentiae.

The Quadroppiidae are a family of oppioid oribatid mites with relatively elongate costulae on the prodorsum.

Characters (from Norton & Behan-Pelletier 2009): Length 150–230 µm. Chelicerae with dentate chelae. Costulae longer than half length of prodorsum, without anterior cusp. Notogaster with nine pairs of setae, all similar in size and shape; crista present, one-third length of notogaster or longer. Borders of epimera III and IV meeting medially; epimeral border IV present; coxisternal setae setose. Solenidion ω2 on tarsus I arising close to famulus; ω1 on tarsus II positioned medially on segment.

<==Quadroppiidae [Quadroppiinae] S04
    |--Borhidia Balogh & Mahunka 1974 [Borhidiidae] S04
    |    |--*B. cubana Balogh & Mahunka 1974 S04
    |    `--B. andina Balogh 1988 S04
    `--Quadroppia Jacot 1939 S04
         |--Q. (Quadroppia) S04
         |    |--*Q. (Q.) quadricarinata (Michael 1885) (see below for synonymy) S04
         |    |--Q. (Q.) cristata Balogh & Mahunka 1980 S04
         |    |--Q. (Q.) hammerae Mínguez, Ruiz & Subías 1985 S04
         |    |--Q. (Q.) illinoisensis (Jacot 1938) [=Oppia quadricarinata illinoisensis] S04
         |    |--Q. (Q.) longisetosa Mínguez, Ruiz & Subías 1985 S04
         |    |--Q. (Q.) maritalis Lions 1982 [=Q. quadricarinata maritalis] S04
         |    `--Q. (Q.) obsoleta Mínguez, Ruiz & Subías 1985 [=Q. longisetosa obsoleta] S04
         `--Q. (Coronoquadroppia Ohkubo 1995) S04
              |--Q. (C.) circumita (Hammer 1961) (see below for synonymy) S04
              |--Q. (C.) abchasica Gordeeva & Tarba 1990 S04
              |--Q. (C.) crenata Mahunka 1984 S04
              |--Q. (C.) expansa (Ohkubo 1995) [=Coronoquadroppia expansa] S04
              |--Q. (C.) galaica Mínguez, Ruiz & Subías 1985 (see below for synonymy) S04
              |--Q. (C.) media Gordeeva 1983 S04
              |--Q. (C.) michaeli Mahunka 1977 [incl. Q. gumista Gordeeva & Tarba 1990, Q. lesleyae Monson 2000] S04
              |--Q. (C.) monstruosa Hammer 1979 (see below for synonymy) S04
              |--Q. (C.) nana Gordeeva 1983 S04
              |--Q. (C.) nasalis Gordeeva 1983 S04
              |--Q. (C.) omodeoi Mahunka & Paoletti 1984 S04
              `--Q. (C.) pseudocircumita Mínguez, Ruiz & Subías 1985 S04

Quadroppia (Coronoquadroppia) circumita (Hammer 1961) [=Oppia circumita; incl. O. quadricarinata ferrumequina Jacot 1938 (n. d.), *Coronoquadroppia parallela Ohkubo 1995] S04

Quadroppia (Coronoquadroppia) galaica Mínguez, Ruiz & Subías 1985 [=Q. pseudocircumita galaica; incl. Q. bellula Luxton 1987] S04

Quadroppia (Coronoquadroppia) monstruosa Hammer 1979 [incl. Q. mahunkai Mínguez 1981, Q. paolii Woas 1986, Q. ritza Gordeeva & Tarba 1990] S04

*Quadroppia (Quadroppia) quadricarinata (Michael 1885) [=Notaspis quadricarinata; incl. Q. skookumchucki Jacot 1939, Q. quadricarinata virginalis Lions 1982] S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Teratoppiidae

Dorsal (A), lateral (B) and ventral (C) views and leg I (D) of Teratoppia centroamericana, from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

The Teratoppiidae are a near-pantropical group (absent in Australia) of oribatid mites with characteristic expanded tibiae bearing lateral apophyses.

Characters (from Norton & Behan-Pelletier 2009): Length 300–450 µm. Gnathosoma normale; chelicerae with dentate chelae, two setae; rutella normally developed. Prodorsum without true lamellae. Notogaster without scalps; octotaxic system absent; dorsophragamata and pleurophragmata absent; paired opisthonotal glands opening directly on notogaster. Epimera III–IV not distinctly delineated, border of epimere III not visible. Genital plates with six pairs of setae. Tibiae I–II expanded laterally, with apophyses; tarsi I–II longer than tibiae I–II; proral setae of tarsi setiform.

<==Teratoppiidae
    |--Granuloteratoppia Balogh 1988 S04
    |    `--*G. annulata Balogh 1988 S04
    |--Leoppia Pérez-Íñigo 1983 S04
    |    `--*L. longicoma Pérez-Íñigo 1983 S04
    |--Brasiloppia Pérez-Íñigo & Baggio 1986 S04
    |    `--*B. flechtmanni Pérez-Íñigo & Baggio 1986 S04
    `--Teratoppia Balogh 1959 S04
         |--T. (Teratoppia) [incl. Cretoppia Mahunka 1986] S04
         |    |--*T. (T.) calcarata Balogh 1959 S04
         |    |--T. (T.) brasiliensis Pérez-Íñigo & Baggio 1996 S04
         |    |--T. (T.) ciliata Wallwork 1961 S04
         |    |--T. (T.) creta (Mahunka 1986) [=Cretoppia creta] S04
         |    |--T. (T.) minor Wallwork 1961 S04
         |    |--T. (T.) reducta Balogh & Mahunka 1969 S04
         |    |--T. (T.) translamellata Mahunka 1983 S04
         |    `--T. (T.) uspiensis Pérez-Íñigo & Baggio 1980 S04
         `--T. (Teratoppiella Balogh 1983) S04
              |--T. (*T.) brevipectinata Balogh & Mahunka 1978 S04
              |--T. (T.) ardua (Pérez-Íñigo & Baggio 1997) [=Teratoppiella ardua] S04
              |--T. (T.) centroamericana Woas 1986 S04
              |--T. (T.) fimbriata (Mahunka 1983) [=Teratoppiella fimbriata] S04
              |--T. (T.) pectinata Balogh 1961 S04
              |--T. (T.) pluripectinata Balogh & Mahunka 1978 (see below for synonymy) S04
              `--T. (T.) regalis Mahunka 1983 S04

Teratoppia (Teratoppiella) pluripectinata Balogh & Mahunka 1978 [incl. T. baloghi Pérez-Íñigo & Baggio 1980, T. pectinata Balogh & Mahunka 1969 non Balogh 1961] S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Ctenobelba

Dorsal (left) and ventral views of Ctenobelba perezinigoi, from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

Ctenobelba is a Palaearctic genus of oribatid mites with tactile first legs.

Characters (from Norton & Behan-Pelletier 2009): Length 400–500 µm. Gnathosoma normal; chelicerae with dentate chelae, two setae; rutella normally developed. Prodorsum without true lamellae, with parallel costulae; shorter in length than notogaster. Notogaster without scalps; with ten pairs of setae, one seta anterior to lyrifissure ia; octotaxic system absent; dorsophragmata and pleurophragmata absent; paired opisthonotal glands opening directly on notogaster. Pedotectum II present. Proral setae of tarsi spiniform.

<==Ctenobelba Balogh 1943 [incl. Elapheremaeus Grandjean 1944; Ctenobelbidae] S04
    |--*C. pectinigera (Berlese 1908) S04 (see below for synonymy)
    |--C. apatomorpha Iturrondobeitia, Saloña et al. 1998 S04
    |--C. brevipilosa Mahunka 1965 S04
    |--C. csiszarae Mahunka 1977 S04
    |--C. foliata Hammer 1961 S04
    |--C. grancanariae Pérez-Íñigo & Peña 1997 S04
    |--C. longisetosa Suzuoka & Aoki 1980 S04
    |--C. mahnerti Mahunka 1974 S04
    |--C. marcuzzii Mahunka 1974 S04
    |--C. obsoleta (Koch 1841) (n. d.) [=Murcia obsoleta] S04
    |--C. parafoliata Pérez-Íñigo 1991 S04
    |--C. perezinigoi Moraza 1985 S04
    |--C. pilosella Jeleva 1962 [incl. C. tuberculata Kulijev 1966] S04
    |--C. polysetosa Aoki & Yamamoto 2000 S04
    |--C. pulchellula Gil-Martín & Subías 1997 S04
    |--C. serrata Mahunak 1964 S04
    |--C. simplex (Willmann 1940) [=Eremobelba simplex] S04
    |--C. soloduchi Pankov 1988 S04
    `--C. translamellata Iorkansky 1990 S04

*Ctenobelba pectinigera (Berlese 1908) S04 [=Eremobelba pectinigera S04, Elapheremaeus pectinigera G46; incl. Amerobelba bicarinata Berlese 1910 S04, Ctenobelba fenestrata Pérez-Íñigo 1990 S04, Notaspis maculosa Warburton & Pearce 1905 (n. d.) S04]

*Type species of generic name indicated

REFERENCES

[G46] Grandjean, F. 1946. Les poils et les organes sensitifs portés par les pattes et le palpe chez les oribates. Bulletin de la Société Zoologique de France 71: 10–29.

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Heterobelbidae

Dorsal view (A), cerotegument (B), leg II (C) and ventral view (D) of Heterobelba barbata, from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

The Heterobelbidae are a mostly pantropical (absent from Australia) group of oribatid mites which carry scalps as adults attached to a pair of humeral tubercles.

Characters (from Norton & Behan-Pelletier 2009): Length 300–400 µm. Gnathosoma normal; chelicerae with dentate chelae, two setae; rutella normally developed. Prodorsum without true lamellae. Scalps present, tightly affixed on pair of humeral tubercles at anterior of notogaster; octotaxic systema absent; dorsophragmata and pleurophragmata absent; paired opisthonotal glands opening directly on notogaster. Genital plates with seven pairs of setae. Tibiae and tarsi without porose areas.

<==Heterobelbidae
    |--Furcodamaeus Pérez-Íñigo & Baggio 1980 S04
    |    `--*F. bifurcatus Pérez-Íñigo & Baggio 1980 S04
    |--Haplobelba Balogh & Mahunka 1969 S04
    |    `--*H. simplex Balogh & Mahunka 1969 S04
    `--Heterobelba Berlese 1913 S04
         |--*H. galerulata Berlese 1913 S04
         |--H. africana Balogh 1958 S04
         |--H. barbata Beck 1962 S04
         |--H. crassisetosa Beck 1962 S04
         |--H. furcata Mahunka 1978 S04
         |--H. magna Beck 1962 S04
         |--H. oxapampensis Beck 1962 S04
         |--H. rostrata Mondal & Kundu 1984 S04
         |--H. spinitecta Mahunka 1983 S04
         |--H. spinosissima Balogh & Mahunka 1981 S04
         |--H. spumosa Mahunka 1983 S04
         |--H. stellifera Okayama 1980 S04
         |    |--H. s. stellifera S04
         |    `--H. s. formosana Aoki 1990 S04
         `--H. zikani Sellnick 1922 S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Eremulidae

Eremulus sp., copyright Biodiversity Institute of Ontario.

Belongs within: Brachypylina.

The Eremulidae are a group of oribatid mites most diverse in tropical regions though some species of Eremulus are also found in temperate habitats. Most genera have three pairs of aggenital setae but five are present in the Argentinian Pseuderemulus gladiator and seven in the palaeotropical Caveremulus.

Characters (from Norton & Behan-Pelletier 2009): Length 275–500 µm. Gnathosoma normal; chelicerae with dentate chelae, two setae; rutella normally developed. Prodorsum shorter in length than notogaster; true lamellae absent, costulae present. Notogaster with scalps absent; octotaxic system absent; dorsophragamata and pleurophragmata absent. paired opisthonotal glands opening direct on notogaster. Pedotectum I present; propodolateral apophysis absent. Ventral plate setae branched; three pairs of adanal setae. Femora I–IV and trochanters III–IV with porose areas; tibiae and tarsi without retrotecta.

Eremulidae
    |--Austroeremulus Mahunka 1985 S04
    |    `--*A. glabrus Mahunka 1985 S04
    |--Pseuderemulus Balogh & Mahunka 1968 S04
    |    `--*P. gladiator Balogh & Mahunka 1968 S04
    |--Reteremuloides Mahunka 1989 S04
    |    `--*R. bifurcatus Mahunka 1989 S04
    |--Caveremulus Mahunka 1983 [incl. Berndbella Mahunka 1985] S04
    |    |--*C. cordisetus Mahunka 1983 S04
    |    `--C. serratus (Mahunka 1985) [=Berndbella serrata] S04
    |--Fenestrella Mahunka 1987 S04
    |    |--*F. bifurcata Mahunka 1987 S04
    |    `--F. sinica Zhao & Wen 1994 S04
    |--Reteremulus Balogh & Mahunka 1966 S04
    |    `--*R. aciculatus Balogh & Mahunka 1966 S04
    |         |--R. a. aciculatus S04
    |         `--R. a. papuanus Balogh 1968 S04
    `--Eremulus Berlese 1908 S04
         |--*E. flagellifer Berlese 1908 [incl. E. modestus Berlese 1908] S04
         |--E. adami Balogh & Mahunka 1966 S04
         |--E. africanus Balogh 1958 (n. d.) S04
         |--E. antis Poltavskaja 1987 S04
         |--E. australis Wen 1988 S04
         |--E. avenifer Berlese 1913 S04
         |--E. baliensis Hammer 1982 S04
         |--E. berlesei Mahunka 1977 S04
         |--E. brasiliensis Pérez-Íñigo & Baggio 1985 S04
         |--E. cingulatus Jacot 1937 S04
         |--E. clavatoflagellatus Lombardini 1963 (n. d.) S04
         |--E. crispus Hammer 1958 S04
         |--E. curviseta Hammer 1971 S04
         |--E. densus Hammer 1979 S04
         |--E. elegans Gordeeva 1987 S04
         |--E. foveolatus Hammer 1962 S04
         |--E. hastatus Hammer 1961 S04
         |--E. lanceocrinus Balogh 1958 (n. d.) S04
         |--E. monstrosus Hammer 1972 S04
         |--E. nigrisetosus Hammer 1958 S04
         |--E. pectinatus Jacot 1937 S04
         |--E. pilipinus Corpuz-Raros 1979 S04
         |--E. renukae Sanyal 1992 S04
         |--E. rigidisetus Balogh & Mahunka 1969 S04
         |--E. rimosus Tseng 1982 S04
         |--E. serratus Hammer 1966 S04
         |--E. spinifer Woolley & Higgins 1963 S04
         |--E. tenuis Hammer 1979 S04
         |--E. translamellatus Balogh & Mahunka 1969 S04
         |--E. truncatus Hammer 1971 S04
         `--E. wallworki Balogh 1958 H07

*Type species of generic name indicated

REFERENCES

[H07] Haq, M. A. 2007. Biodegradation and productivity in tropical ecosystem by oribatid mites. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 137–150. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 24 February 2019.

Basilobelbidae

Dorsal and ventral views of Basilobelba sollertius, from Norton & Behan-Pelletier (2009).

Belongs within: Brachypylina.

The Basilobelbidae are a pantropical group of oribatid mites bearing scalps attached to the notogaster by an anterior medial tubercle. The chelicerae are peloptoid and covered by a pointed rostrum in the genus Xiphobelba whereas Basilobelba has dentate chelicerae and a rounded rostrum.

Characters (from Norton & Behan-Pelletier 2009): Length 400–750 µm. Gnathosoma normal; chelicerae with dentate chelae, two setae; rutella normally developed. Prodorsum without true lamellae; bothridial seta with ringlike thickenings proximally. Scalps present, tightly affixed on medial tubercle at anterior of notogaster; octotaxic system absent; dorsophragmata and pleurophragmata absent; paired opisthonotal glands opening directly on notogaster. Circumventral carina present curving posterior to anal plates; genital plates with seven pairs of setae. Trochanter II with broad ventral keel mimicking pedotectum II; tibiae and tarsi without porose areas; setae p absent from tarsi II to IV.

Basilobelbidae
    |--Xiphobelba Csiszár 1961 S04
    |    |--*X. hamanni Csiszár 1961 [incl. X. margosetosa Hammer 1979] S04
    |    |--X. barbata Talukdar & Chakrabarti 1984 S04
    |    |--X. ismalia Haq 1980 S04
    |    |--X. mindanensis Corpuz-Raros 1980 S04
    |    `--X. setosa Aoki 1968 S04
    `--Basilobelba Balogh 1958 [incl. Hammation Grandjean 1959] S04
         |--*B. retiaria (Warburton 1912) S04 [=Damaeus retiarius S04, B. retiarius CH98, Hammation retiarius H98]
         |    |--B. r. retiarius  S04
         |    `--B. r. symmetrica Wallwork 1962 S04
         |--B. aethiopica Bernini 1988 S04
         |--B. africana Wallwork 1961 S04
         |--B. atimonensis Corpuz-Raros 1980 S04
         |--B. baltazarae Corpuz-Raros 1979 S04
         |--B. barbata Mondal & Kundu 1984 S04
         |--B. cendanai Corpuz-Raros 1980 S04
         |--B. foliata Hammer 1982 S04
         |--B. indica Bhaduri, Chakrabarti & Raychaudhuri 1974 S04
         |--B. insularis Mahunka 1985 S04
         |--B. monstruosetosa Mahunka 1974 S04
         |--B. pacifica Hammer 1971 S04
         |--B. parmata Okayama 1980 S04
         |--B. sollertius (Grandjean 1959) NB-P09, S04 [=*Hammation sollertius CH98]
         `--B. werneri Mahunka 1982 S04

*Type species of generic name indicated

REFERENCES

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W. & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 22 December 2018.

Polypterozetoidea

Podopterotegaeus tectus, from Canadian Biodiversity Information Facility.

Belongs within: Circumdehiscentiae.

The Polypterozetoidea is a relatively small group of disparate oribatid mites found in cool habitats in temperate regions or at high elevations (Norton & Behan-Pelletier 2009).

Characters (from Norton & Behan-Pelletier 2009): Adults with scalps present or absent. Prodorsum with broad lamellae, translamella present or absent, without genal notch. Tutorium present. Pedotectum I present; patronium present or absent; pedotectum II present or absent. Dorsophragmata and pleurophragmata absent. Subcapitulum diarthric or anarthric, with modified chelicerae and rutella; subcapitular membrane with tectum; palpal eupathidium acm separate from recumbent solenidion. Notogaster often obscured by dirt adhered to cerotegument; with humeral apophysis that may resemble pteromorph; with or without medial apophysis. Six pairs of genital setae. Immature of Polypterozetes cherubin eupheredermous; hysterosoma usually obscured by dirt adhered to cerotegument; quadrideficient; seta d present to tritonymph on tibiae and genua when respective solenidia present (immatures of other species unknown).

<==Polypterozetoidea S04
    |--Polypterozetes Berlese 1916 [Polypterozetidae] S04
    |    `--*P. cherubin Berlese 1916 BB92
    |--Podopterotegaeus Aoki 1969 [Podopterotegaeidae] S04
    |    |--*P. tectus Aoki 1969 BB92
    |    |--P. altimonticola Piffl 1972 S04
    |    `--P. bisetus Pankov 1984 S04
    |--Tumerozetes Hammer 1966 S04 [Tumerozetidae NB-P09]
    |    |--*T. bifurcatus Hammer 1966 BB92
    |    |--T. circularis Hammer 1966 S04
    |    |--T. indistinctus Hammer 1966 S04
    |    |--T. parallelus Hammer 1966 S04
    |    `--T. pumilis Hammer 1966 S04
    `--Nodocepheidae NB-P09
         |--Nemacepheus Aoki 1968 S04
         |    `--*N. dentatus Aoki 1968 S04
         `--Nodocepheus Hammer 1958 S04
              |--*N. dentatus Hammer 1958 S04
              |    |--N. d. dentatus S04
              |    `--N. d. barbatus Hammer 1966 S04
              |--N. baloghi Mahunka 1983 S04
              |--N. cerebralis Mahunka 1980 S04
              |--N. hammerae Balogh 1961 S04
              |--N. laterodentatus Piffl 1972 S04
              `--N. minimus Mahunka 1985 S04

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Atropacarus

Atropacarus sp., copyright Scott Justis.

Belongs within: Phthiracaridae.

Atropacarus is a cosmopolitan genus of phthiracarid mites with three or four pairs of closely-placed setae on the inner margin of the ano-adanal plates. The type subgenus Atropacarus, with four pairs of such setae, is sometimes treated separately from the remaining subgenera with three pairs, which are then placed as the genus Hoplophorella. Members of the subgenus Kakophthiracarus are characterised by having all setae in the anogenital region very short and fine.

<==Atropacarus Ewing 1917 [incl. Helvetacarus Mahunka 1993] S04
    |--A. (Atropacarus) N07
    |    |--*A. (A.) striculus (Koch 1835) S04 [=Hoplophora stricula S04, Steganacarus striculus G34]
    |    |    |--A. s. striculus (see below for synonymy) S04
    |    |    `--A. s. insularis (Weigmann 1976) [=Steganacarus insularis] S04
    |    |--A. (A.) absimilis (Niedbała 1982) [=Steganacarus (A.) absimilis] S04
    |    |--A. (A.) clavatus Aoki 1980 N86a [=A. striculus clavatus A80]
    |    |--A. (A.) clavigerus (Berlese 1904) N84 (see below for synonymy)
    |    |--A. (A.) csiszarae (Balogh & Mahunka 1979) [=Steganacarus csiszarae] N86a
    |    |--A. (A.) decipiens (Niedbała 1984) [=Steganacarus (A.) decipiens] S04
    |    |--A. (A.) echinodiscus (Mahunka 1982) [=Steganacarus echinodiscus] N86a
    |    |--A. (A.) folious Niedbała 2003 N07
    |    |--A. (A.) griseus (Niedbała 1984) [=Steganacarus (A.) griseus] S04
    |    |--A. (A.) immundus (Niedbała 1983) [=Steganacarus (A.) immundus] S04
    |    |--A. (A.) inculpatus (Niedbała 1984) [=Steganacarus (A.) inculpatus] S04
    |    |--A. (A.) maculosus (Niedbała 1983) [=Steganacarus (A.) maculosus] S04
    |    |--A. (A.) mirabilis (Mahunka 1979) [=Steganacarus mirabilis] N86a
    |    |--A. (A.) murciensis Niedbała 1986 N86b
    |    |--A. (A.) obesus (Niedbała 1983) [=Steganacarus (A.) obesus] S04
    |    |--A. (A.) ochraceus (Niedbała 1983) [=Steganacarus (A.) ochraceus] S04
    |    |--A. (A.) parvulus (Niedbała 1983) [=Steganacarus (A.) parvulus] S04
    |    |--A. (A.) perversus (Niedbała 1983) [=Steganacarus (A.) perversus] S04
    |    |--A. (A.) phyllophorus (Berlese 1904) N86a (see below for synonymy)
    |    |--A. (A.) platakisi (Mahunka 1979) N86a, S04 (see below for synonymy)
    |    |--A. (A.) serratus (Feider & Suciu 1957) N84, S04 [=Steganacarus serratus S04]
    |    |--A. (A.) substrictus (Niedbała 1983) [=Steganacarus (A.) substrictus] S04
    |    |--A. (A.) terrapene (Jacot 1937) [=Steganacarus terrapene] N86a
    |    `--A. (A.) wandae (Niedbała 1981) N86a [=Steganacarus wandae S04; incl. S. navarrensis Moraza 1985 S04]
    |--A. (Hoplophorella Berlese 1923) N07, S04
    |    |--A. (*H.) cucullatus (Ewing 1909) BB92, S04 (see below for synonymy)
    |    |    |--A. c. cucullatus (see below for synonymy) S04
    |    |    `--‘Hoplophorella’ c. curassensis Willmann 1936 S04
    |    |--‘Hoplophorella’ (H.) angolensis Mahunka 1984 S04
    |    |--‘Hoplophorella’ (H.) berninii (Niedbała 1991) [=Protophthiracarus berninii] S04
    |    |--‘Hoplophorella’ (H.) brasiliensis (Niedbała 1988) [=Austrophthiracarus brasiliensis] S04
    |    |--A. (H.) brevipilis (Balogh 1958) [=Steganacarus brevipilis] N86a
    |    |--‘Hoplophorella’ (H.) collaris (Balogh 1958) [=Steganacarus collaris] S04
    |    |--‘Hoplophorella’ (H.) craterifer (Hammer 1971) S04 (see below for synonymy)
    |    |--A. (H.) diaphoros Niedbała 2000 S04
    |    |--‘Hoplophorella’ (H.) digna Niedbała 1983 S04 [=Protophthiracarus dignus N86a]
    |    |--A. (H.) dissimilis Niedbała 1998 S04
    |    |--‘Hoplophorella’ (H.) dubia (Niedbała 1992) S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) ensifera Mahunka 1985 S04
    |    |--‘Hoplophorella’ (H.) ephippigera (Balogh & Mahunka 1977) [=Steganacarus ephippiger] S04
    |    |--‘Hoplophorella’ (H.) evergladensis (Niedbała 2001) [=Protophthiracarus evergladensis] S04
    |    |--‘Hoplophorella’ (H.) fonseciai (Pérez-Íñigo & Baggio 1980) S04 (see below for synonymy)
    |    |--A. (H.) frondeus Niedbała 2003 N07
    |    |--‘Hoplophorella’ (H.) grandjeani Niedbała 1981 S04 [=Protophthiracarus grandjeani N86a]
    |    |--A. (H.) hamatus (Ewing 1909) N07 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) heterotricha Mahunka 1984 S04
    |    |--A. (H.) insolens (Neidbala 1988) N86a, S04 [=Hoplophorella insolens S04]
    |    |--A. (H.) kulczynskii (Niedbała 1981) [=Hoplophorella kulczynskii] N86a
    |    |--‘Hoplophorella’ (H.) lanceoseta (Balogh & Mahunka 1981) S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) lanceosetoides Mahunka 1985 S04
    |    |--‘Hoplophorella’ (H.) lemuria Mahunka 1993 S04
    |    |--‘Hoplophorella’ (H.) manipurensis Misra, Bhaduri & Raychaudhuri 1982 S04
    |    |--A. (H.) minisetosus (Mahunka 1978) [=Hoplophorella minisetosa] N86a
    |    |--‘Hoploderma’ multipunctatum Sellnick 1918 S93
    |    |--‘Hoplophorella’ (H.) neonominata Subías 2004 (see below for synonymy) S04
    |    |--A. (H.) nigeriensis Badejo, Woas & Beck 2001 S04
    |    |--A. (H.) oblongus (Niedbała 1983) [=Hoplophorella oblonga] N86a
    |    |--A. (H.) pallens (Niedbała 1983) N91, S04 [=Hoplophorella pallens S04]
    |    |--‘Hoplophorella’ (H.) paraensis (Pérez-Íñigo & Baggio 1996) [=Protophthiracarus paraensis] S04
    |    |--A. (H.) pergratus Niedbała 1998 [=Hoplophorella pergrata] S04
    |    |--‘Hoplophorella’ (H.) perisi Subías & Sarkar 1984 S04
    |    |--‘Hoplophorella’ (H.) pervicax (Niedbała 1984) S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) praeoccupata Subías 2004 (see below for synonymy) S04
    |    |--‘Hoplophorella’ (H.) prominens (Balogh 1958) S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) pustulata Mahunka 1993 S04
    |    |--A. (H.) rangiroaensis (Hammer 1972) [=Hoplophorella rangiroaensis] N86a
    |    |--‘Hoplophorella’ (H.) remigera (Berlese 1923) [=Phthiracarus (Trachyhoplophora) remiger] S04
    |    |--‘Hoplophorella’ (H.) repetita Subías 2004 (see below for synonymy) S04
    |    |--‘Hoplophorella’ (H.) sabahna Mahunka 1991 S04
    |    |--‘Hoplophorella’ (H.) sculptilis (Niedbała 1988) S04 (see below for synonymy)
    |    |--A. (H.) singularis (Sellnick 1959) N91, S04 (see below for synonymy)
    |    |--A. (H.) stilifer (Hammer 1961) N91, S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) subciliata Mahunka 1983 S04
    |    |--A. (H.) subitus (Niedbała 1983) [=Hoplophorella subita] N86a
    |    |--‘Hoplophorella’ (H.) subsellata Balogh & Mahunka 1977 S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (H.) tripartita (Niedbała & Schatz 1996) [=Protophthiracarus tripartitus] S04
    |    |--‘Hoplophorella’ (H.) unqus (Niedbała 1991) [=Notophthiracarus unqus] S04
    |    |--‘Hoplophorella’ (H.) varia (Niedbała & Schatz 1996) [=Protophthiracarus varius] S04
    |    |--‘Hoplophorella’ (H.) varians Jacot 1933 S04
    |    |--‘Hoplophorella’ (H.) venusta Niedbała 1983 S04 [=Protophthiracarus venustus N86a]
    |    `--A. (H.) vitrinus (Berlese 1913) N01, S04 (see below for synonymy)
    |--‘Hoplophorella’ (Kakophthiracarus Mahunka 1992) S04
    |    |--A. (K.) tuberculosissima (Mahunka 1978) N86a, S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (K.) balazsi Mahunka 1983 S04
    |    |--‘Hoplophorella’ (K.) cavernosa Mahunka 1987 S04
    |    |--‘Hoplophorella’ (K.) enigmatica (Mahunka 1996) [=Kakophthiracarus enigmaticus] S04
    |    |--A. (K.) multirugosus (Mahunka 1978) N86a, S04 (see below for synonymy)
    |    |--‘Hoplophorella’ (K.) multituberculata (Balogh & Mahunka 1966) S04 (see below for synonymy)
    |    `--‘Hoplophorella’ (K.) mwali (Mahunka 1994) [=Kakophthiracarus mwali] S04
    `--‘Hoplophorella’ (Steganacarellus Mahunka 1986) S04
         |--‘Hoplophorella’ (*S.) reducta (Mahunka 1986) [=*Steganacarellus reductus] S04
         |--‘Hoplophorella’ (S.) heterosetosa (Niedbała 1988) [=Notophthiracarus heterosetosus] S04
         |--‘Hoplophorella’ (S.) ‘heterotricha’ (Niedbała 2000) (see below for synonymy) S04
         |--‘Hoplophorella’ (S.) paraparvula (Niedbała 2000) [=Notophthiracarus paraparvulus] S04
         |--‘Hoplophorella’ (S.) parvula (Niedbała 1998) [=Notophthiracarus parvulus] S04
         `--‘Hoplophorella’ (S.) perlucunda (Niedbała 2000) [=Notophthiracarus perlucundus] S04

Atropacarus incertae sedis:
  A. bichei Niedbała 1986 S04
  A. ciliosus Niedbała 2002 S04
  A. clavatus Aoki 1980 [=A. sticulus clavatus] S04
  A. controversus Niedbała 2000 S04
  A. genavensis (Mahunka 1993) [=Helvetacarus genavensis] S04
  A. inconditus Mahunka 1991 S04
  A. macrosculpturatus (Mahunka & Mahunka-Papp 1999) [=Steganacarus macrosculpturatus] S04
  A. multisetosus Niedbała 2002 S04
  A. paraclavatus Niedbała 2002 S04
  A. punctulatus Moraza 1985 S04
  ‘Hoplophorella’ thoreaui (Jacot 1930) N83

Atropacarus (Atropacarus) clavigerus (Berlese 1904) N84 [=Hoploderma clavigerum N84, Steganacarus (A.) clavigerus N84; incl. Phthiracarus lutulentus Sellnick 1920 S04]

Atropacarus (Atropacarus) phyllophorus (Berlese 1904) N86a [=Hoploderma phyllophorum S04, Steganacarus phyllophorus N86a; incl. S. ropalus Feider & Suciu 1957 S04]

Atropacarus (Atropacarus) platakisi (Mahunka 1979) N86a, S04 [=Steganacarus platakisi S04; incl. A. papillatus Gil-Martín, Subías & Arillo 1992 S04]

*Atropacarus (Atropacarus) striculus striculus (Koch 1835) [incl. Steganacarus diaphanus Jacot 1930, S. senex Aoki 1958, S. trichosus Mahunka 1982] S04

Atropacarus (*Hoplophorella) cucullatus (Ewing 1909) BB92, S04 [=Hoploderma cucullatum S04, Hoplophora cucullatum BB92, *Hoplophorella cucullata S04]

Atropacarus (*Hoplophorella) cucullatus cucullatus (Ewing 1909) [incl. Hoplophorella cucullata cuculloides Jacot 1933, Hoploderma licnophorum Berlese 1913] S04

Atropacarus (Hoplophorella) hamatus (Ewing 1909) N07 [=Hoploderma hamatum S04, Hoplophorella hamata S04; incl. Hoplophorella cochlearia Pérez-Íñigo & Baggio 1993 S04, Hoplophorella cuneiseta Mahunka 1988 S04, Hoplophorella cucullata floridae Jacot 1933 S04, Atropacarus (Hoplophorella) floridae N91, Hoplophorella glauca Hammer 1972 S04, A. (H.) glaucus N07, Hoplophorella ligulifera Mahunka 1987 S04, Phthiracarus (H.) cucullatus obsoletior Berlese 1923 S04, Hoplophorella schauenbergi Mahunka 1978 S04, Atropacarus (H.) schauenbergi N91, Hoplophorella spatulata Parry 1980 S04]

Atropacarus (Hoplophorella) singularis (Sellnick 1959) N91, S04 [=Hoplophorella singularis S04; incl. H. queenslandicus Balogh & Mahunka 1978 S04, H. regalis Mahunka 1978 S04, H. sundarbanensis Sanyal & Bhaduri 1982 S04]

Atropacarus (Hoplophorella) stilifer (Hammer 1961) N91, S04 [=Steganacarus stilifer S04, Hoplophorella stilifera S04; incl. Hoplophorella benoiti Mahunka 1984 S04]

Atropacarus (Hoplophorella) vitrinus (Berlese 1913) N01, S04 [=Hoploderma vitrinum S04, Hoplophorella vitrina S04; incl. Hoplophorella africana Wallwork 1967 S04, Steganacarus andrei Balogh 1958 S04, Hoplophorella lienhardi Mahunka 1987 S04, H. raychaudhurii Subías & Sarkar 1984 S04, H. scapellata Aoki 1965 S04, Atropacarus (Hoplophorella) scapellatus N91]

Atropacarus (Kakophthiracarus) multirugosus (Mahunka 1978) N86a, S04 [=Steganacarus multirugosus S04, Hoplophorella (K.) multirugosa S04; incl. S. cornutus Mahunka 1978 S04]

Atropacarus (Kakophthiracarus) tuberculosissimus (Mahunka 1978) N86a, S04 [=Steganacarus tuberculosissimus S04, Hoplophorella (K.) tuberculosissima S04; incl. H. horrida Mahunka 1984 S04, H. meszarosi Mahunka 1988 S04, H. (*Kakophthiracarus) tuberosa Mahunka 1988 S04, H. verrucosa Mahunka 1987 S04]

‘Hoplophorella’ (Hoplophorella) craterifer (Hammer 1971) S04 [=Steganacarus craterifer S04, Notophthiracarus craterifer N86a]

‘Hoplophorella’ (Hoplophorella) dubia (Niedbała 1992) S04 [=Hoplophthiracarus dubius S04, Austrophthiracarus dubius N86a]

‘Hoplophorella’ (Hoplophorella) fonseciai (Pérez-Íñigo & Baggio 1980) S04 [=Steganacarus fonseciai S04, Protophthiracarus fonseciai N86a]

‘Hoplophorella’ (Hoplophorella) lanceoseta (Balogh & Mahunka 1981) S04 [=Steganacarus lanceosetus S04; incl. H. neglecta Niedbała 1984 S04, Atropacarus (Hoplophorella) neglectus N86a]

‘Hoplophorella’ (Hoplophorella) neonominata Subías 2004 [=Protophthiracarus cavernosus Niedbała 2002 non H. cavernosa Mahunka 1987] S04

‘Hoplophorella’ (Hoplophorella) pervicax (Niedbała 1984) S04 [=Hoplophthiracarus pervicax S04, Austrophthiracarus pervicax N86a]

‘Hoplophorella’ (Hoplophorella) praeoccupata Subías 2004 [=Atropacarus (H.) tuberosus Niedbała & Schatz 1996 non H. tuberosa Mahunka 1988] S04

‘Hoplophorella’ (Hoplophorella) prominens (Balogh 1958) S04 [=Steganacarus prominens S04, Protophthiracarus promineus N86a]

‘Hoplophorella’ (Hoplophorella) repetita Subías 2004 [=Protophthiracarus reductus Niedbała 1991 nec Steganacarellus reductus Mahunka 1986 nec Hoplophorella reducta Mahunka 1988] S04

‘Hoplophorella’ (Hoplophorella) sculptilis (Niedbała 1988) S04 [=Hoplophthiracarus sculptilis S04, Austrophthiracarus sculptilis N86a; incl. Hoplophthiracarus magnus Mahunka 1988 S04]

‘Hoplophorella’ (Hoplophorella) subsellata Balogh & Mahunka 1977 S04 [=Protophthiracarus subsellatus N86a; incl. Steganacarus galeatus Balogh & Mahunka 1978 S04]

‘Hoplophorella’ (Kakophthiracarus) multituberculata (Balogh & Mahunka 1966) S04 [=Steganacarus multituberculatus S04, Notophthiracarus multituberculatus N86a]

‘Hoplophorella’ (Steganacarellus) ‘heterotricha’ (Niedbała 2000) [=Arphthicarus heterotrichus non H. heterotricha Mahunka 1984] S04

*Type species of generic name indicated

REFERENCES

[A80] Aoki, J. 1980. A revision of the oribatid mites of Japan. I. The families Phthiracaridae and Oribotritiidae. Bulletin of the Institute of Environmental Science and Technology, Yokohama National University 6 (2): 1–89.

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[G34] Grandjean, F. 1934. Phthiracarus anonymum, n. sp. Revue Française d'Entomologie 1 (1): 51–58.

[N83] Niedbała, W. 1983. Phthiracaridae nouveaux de l'Ouganda (Acari, Oribatida). Folia Entomologica Hungarica 44 (1): 109–123.

[N84] Niedbała, W. 1984. Phthiracaridae (Acari, Oribatida) nouveaux d'Asie occidentale. Annales Zoologici 38 (10): 225–241.

[N86a] Niedbała, W. 1986a. Catalogue des Phthiracaroidea (Acari), clef pour la détermination des espèces et descriptions d'espèces nouvelles. Annales Zoologici 40 (4): 309–370.

[N86b] Niedbała, W. 1986b. Phthiracaroidea (Acari Oribatida Euptyctima) nouveaux d'Espagne meridionale. Redia 69: 93–104.

[N91] Niedbała, W. 1991. Preliminary hypotheses on the biogeography of the Phthiracaroidea (Acari, Oribatida). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 219–231. SPB Academic Publishing: The Hague.

[N01] Niedbała, W. 2001. Study of the diversity of Ptyctima (Acari: Oribatida) and quest for centres of its origin: the fauna of the Oriental and Australian regions. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 143–148. CSIRO Publishing: Melbourne.

[N07] Niedbała, W. 2007. Ptyctimous mites (Acari, Oribatida) of Costa Rica. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 39–42. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 7 March 2019.

Notophthiracarus

Notophthiracarus bentoni, copyright Mo Hassan.

Belongs within: Phthiracaridae.

Notophthiracarus is a genus of ptyctimous oribatid mites that is particularly diverse in the Oriental and Australasian regions. Members of the subgenus Notophthiracarus sensu stricto have the genital setae arranged in a single longitudinal row along the inner margin of the genital plates but two such rows are present in Calyptophthiracarus. In members of the subgenus Besuchetacarus, a pair of longitudinal keels run the entire length of the notogaster.

<==Notophthiracarus Ramsay 1966 [Calyptophthiracarini] S04
    |  i. s.: N. pedanos Niedbała 2003 N07
    |--N. (Notophthiracarus) S04
    |    |--*N. (N.) maculatus (Traegårdh 1931) [=Phthiracarus maculatus] S04
    |    |--N. (N.) abstemius Niedbała & Colloff 1997 S04
    |    |--N. (N.) aculeatus Niedbała 1988 S04
    |    |--N. (N.) alienus Niedbała 1989 S04
    |    |--N. (N.) alpinus Balogh & Mahunka 1997 S04
    |    |--N. (N.) aokii (Mahunka 1983) [=Hoplophthiracarus aokii] S04
    |    |--N. (N.) aquilus Niedbała 2000 S04
    |    |--N. (N.) araios (Niedbała 2001) [=Protophthiracarus araios] S04
    |    |--N. (N.) ater Niedbała 2000 S04
    |    |--N. (N.) australis Ramsay 1966 [incl. N. caliginosus Niedbała 1989] S04
    |    |--N. (N.) austroafricanus (Mahunka 1984) [=Hoplophorella austroafricana] S04
    |    |--N. (N.) bentoni Niedbała 1998 S04
    |    |--N. (N.) calugari Niedbała 1987 S04
    |    |--N. (N.) capillatus Niedbała 1989 S04
    |    |--N. (N.) caudatus (Balogh & Mahunka 1977) [=Phthiracarus caudatus] S04
    |    |--N. (N.) claviger Niedbała 1993 S04
    |    |--N. (N.) comparativus Niedbała & Colloff 1997 S04
    |    |--N. (N.) consimilis Niedbała 1989 S04
    |    |--N. (N.) conspicuus Niedbała 1989 S04
    |    |--N. (N.) contortulus Niedbała 1992 S04
    |    |--N. (N.) curiosus Niedbała 1998 S04
    |    |--N. (N.) dactyloscopicus Mahunka 1978 S04 [=Hoplophthiracarus dactyloscopicus N86]
    |    |--N. (N.) distinctus Niedbała 1989 S04 [=N. disiunctus (l. c.) H98]
    |    |--N. (N.) echinus (Balogh 1964) [=Hoplophorella echinus] S04
    |    |--N. (N.) endroedryyoungai (Mahunka 1984) [=Hoplophorella endroedryyoungai] S04
    |    |--N. (N.) evexus (Niedbała 2000) [=Arphthicarus evexus] S04
    |    |--N. (N.) fatidicus (Niedbała 1982) [=Hoplophthiracarus fatidicus] S04
    |    |--N. (N.) fecundus Niedbała 2000 S04
    |    |--N. (N.) feideri (Balogh & Csiszár 1963) [=Phthiracarus feideri] S04
    |    |--N. (N.) flagrus Niedbała 2000 S04
    |    |--N. (N.) flexiloquus Niedbała 1989 S04
    |    |--N. (N.) fornicarius (Niedbała 1982) [=Phthiracarus fornicarius] S04
    |    |--N. (N.) fulvus (Niedbała 1985) [=Hoplophthiracarus fulvus] S04
    |    |--N. (N.) furcatus (Niedbała & Corpuz-Raros 1998) [=Arphthicarus furcatus] S04
    |    |--N. (N.) fusticulus (Niedbała 2000) [=Austrophthiracarus fusticulus] S04
    |    |--N. (N.) gressitti Balogh & Mahunka 1997 S04
    |    |--N. (N.) hamidi (Mahunka 1987) [=Hoplophthiracarus hamidi] S04
    |    |--N. (N.) hammeni Niedbała 1987 S04
    |    |--N. (N.) hammerae Niedbała 1987 S04 [=N. hammeri N87]
    |    |--N. (N.) ignobilis Niedbała 2000 S04
    |    |--N. (N.) improvisus (Niedbała 1982) [=Phthiracarus improvisus] S04
    |    |--N. (N.) inaequus (Niedbała 1984) S04 [=Hoplophthiracarus inaequus S04, Austrophthiracarus inaequus N86]
    |    |--N. (N.) inauditus (Niedbała 1982) [=Phthiracarus inauditus] S04
    |    |--N. (N.) incomparabilis Niedbała 2000 S04
    |    |--N. (N.) indicus (Bayoumi & Mahunka 1979) S04 (see below for synonymy)
    |    |--N. (N.) indiligens (Niedbała 1984) S04 (see below for synonymy)
    |    |--N. (N.) indubiatus Niedbała & Colloff 1997 S04
    |    |--N. (N.) inenarrabilis (Niedbała 1982) S04 (see below for synonymy)
    |    |--N. (N.) ineptus (Niedbała 1984) S04 [=Hoplophthiracarus ineptus S04, Austrophthiracarus ineptus N86]
    |    |--N. (N.) kamillae Niedbała 1987 S04 [=N. kamilli CH98]
    |    |--N. (N.) latior (Niedbała 1982) S04 [=Hoplophthiracarus latior S04, Calyptophthiracarus latior N87]
    |    |--N. (N.) leei Niedbała 1987 S04 [=N. lee N87]
    |    |--N. (N.) lineolatus Mahunka 1993 S04
    |    |--N. (N.) lionsi Niedbała 1987 S04
    |    |--N. (N.) longisetus Niedbała 2000 S04
    |    |--N. (N.) mahunkai Niedbała 1987 S04
    |    |--N. (N.) marginatus (Nahunka 1984) [=Hoplophthiracarus marginatus] S04
    |    |--N. (N.) maurus Niedbała 2000 S04
    |    |--N. (N.) mayottei (Niedbała 2001) [=Protophthiracarus mayottei] S04
    |    |--N. (N.) michaeli (Niedbała 1987) S04 [=Calyptophthiracarus michaeli S04, Austrophthiracarus michaeli CH98]
    |    |--N. (N.) mirus Niedbała 2000 S04
    |    |--N. (N.) modicus Niedbała 2000 S04
    |    |--N. (N.) nicoleti (Niedbała 1987) S04 [=Calyptophthiracarus nicoleti S04, Austrophthiracarus nicoleti CH98]
    |    |--N. (N.) nigerrimus (Berlese 1920) [=Phthiracarus nigerrimus] S04
    |    |--N. (N.) notatus (Niedbała 2001) [=Arphthicarus notatus] S04
    |    |--N. (N.) obsessus Subías 2004 S04 (see below for synonymy)
    |    |--N. (N.) paracuriosus Niedbała 2000 S04
    |    |--N. (N.) parasentus (Niedbała 2000) [=Arphthicarus parasentus] S04
    |    |--N. (N.) parasomalicus Niedbała 2001 S04
    |    |--N. (N.) parasummersi Niedbała 2001 S04
    |    |--N. (N.) paravariolosus Niedbała 1994 S04
    |    |--N. (N.) pardinus (Berlese 1916) [=Hoploderma pardinum] S04
    |    |--N. (N.) parilloi Niedbała 2001 S04
    |    |--N. (N.) pauliani (Balogh 1964) [=Hoplophorella pauliani] S04
    |    |--N. (N.) perezinigoi Niedbała 1987 S04
    |    |--N. (N.) perti (Niedbała 1987) S04 [=Calyptophthiracarus perti S04, Austrophthiracarus perti CH98]
    |    |--N. (N.) procerus Niedbała 2001 S04
    |    |--N. (N.) quietus Niedbała 1989 S04
    |    |--N. (N.) rafalskii (Niedbała 1997) S04 (see below for synonymy)
    |    |--N. (N.) ramsayi Niedbała 1987 S04 [=N. ramsai N87]
    |    |--‘Hoplophorella’ reducta Mahunka 1988 (see below for synonymy) S04
    |    |--N. (N.) remotus (Niedbała 1989) S04 [=Arphthicarus remotus S04, Austrophthiracarus remotus CH98]
    |    |--N. (N.) repostus Niedbała 1989 S04
    |    |--N. (N.) sacyae (Mahunka 1983) [=Steganacarus sacyae] S04
    |    |--N. (N.) schatzi Niedbała 1989 S04
    |    |--N. (N.) schizocomus (Hammer 1962) S04 (see below for synonymy)
    |    |--N. (N.) schusteri Niedbała 1987 S04
    |    |--N. (N.) sentus (Niedbała 1989) [=Austrophthiracarus sentus] S04
    |    |--N. (N.) shealsi (Lee 1981) [=Hoplophthiracarus shealsi] S04
    |    |--N. (N.) sicilicomus (Hammer 1962) [=Phthiracarus sicilicomus] S04
    |    |--N. (N.) similis Niedbała 2001 S04
    |    |--N. (N.) sinuosus (Niedbała 1982) [=Phthiracarus sinuosus] S04
    |    |--N. (N.) solitarius Niedbała & Colloff 1997 S04
    |    |--N. (N.) solomonensis Niedbała 1998 S04
    |    |--N. (N.) somalicus (Berlese 1923) [=Phthiracarus (Trachyhoplophora) somalicus] S04
    |    |--N. (N.) sordidus Niedbała & Colloff 1997 S04
    |    |--N. (N.) sororius (Niedbała 1982) S04 [=Hoplophthiracarus sororius S04, Austrophthiracarus sororius N86]
    |    |--N. (N.) spurcus Niedbała & Colloff 1997 S04
    |    |--N. (N.) stenotus Niedbała & Corpuz-Raros 1998 S04
    |    |--N. (N.) sumatranus (Niedbała 2000) [=Arphthicarus sumatranus] S04
    |    |--N. (N.) summersi Niedbała 2001 S04
    |    |--N. (N.) tinctus (Niedbała 2000) [=Arphthicarus tinctus] S04
    |    |--N. (N.) tohivea Niedbała 1998 S04
    |    |--N. (N.) trichosus (Mahunka 1988) [=Hoplophthiracarus trichosus] S04
    |    |--N. (N.) tripartitus Niedbała 1989 S04
    |    |--N. (N.) uncinatus Niedbała & Colloff 1997 S04
    |    |--N. (N.) uncinulus Niedbała 2000 S04
    |    |--N. (N.) unicarinatus Niedbała 2000 S04
    |    |--N. (N.) usitatus Niedbała 1989 S04
    |    |--N. (N.) variolosus (Berlese 1888) [=Hoplophora variolosa] S04
    |    |--N. (N.) veteratorius (Niedbała 1988) S04 (see below for synonymy)
    |    |--N. (N.) weigmanni Niedbała 1987 S04
    |    `--N. (N.) zebra (Balogh 1964) [=Hoplophorella zebra] S04
    |--N. (Besuchetacarus Mahunka 1985) S04
    |    |--N. (*B.) orientalis (Mahunka 1985) [=*Besuchetacarus orientalis] S04
    |    |--N. (B.) bicarinatus Niedbała 2001 S04
    |    |--N. (B.) hauseri Mahunka 1995 S04
    |    |--N. (B.) lienhardi Mahunka 1996 S04
    |    `--N. (B.) tricarinatus (Niedbała 1988) [=Calyptophthiracarus tricarinatus] S04
    `--N. (Calyptophthiracarus Aoki 1980) [incl. Hauserophthiracarus Mahunka 1982, Phrathicarus Niedbała 1994] S04
         |--N. (*C.) mitratus (Aoki 1980) [=*Calyptophthiracarus mitratus] S04
         |--N. (C.) canariensis (Pérez-Íñigo & Peña 1996) [=Calyptophthiracarus canariensis] S04
         |--N. (C.) candidulus (Niedbała 1983) [=Hoplophthiracarus candidulus] S04
         |--N. (C.) comatus Niedbała 2000 S04
         |--N. (C.) concolor (Sergienko 2000) [=Calyptophthiracarus concolor] S04
         |--N. (C.) costai (Macfarlane & Sheals 1965) [=Hoplophthiracarus costai] S04
         |--N. (C.) excellens (Niedbała 1982) S04 [=Neophthiracarus excellens S04, Austrophthiracarus excellens CH98]
         |--‘Calyptophthiracarus’ globiger (Hammer 1962) [=Phthiracarus globiger] N86
         |--N. (C.) heterotrichus (Mahunka 1979) [=Hoplophthiracarus heterotrichus] S04
         |--N. (C.) inflatus (Niedbała 1994) [=Phrathicarus inflatus] S04
         |--N. (C.) inusitatus (Niedbała 1983) [=Hoplophthiracarus inusitatus] S04
         |--N. (C.) meridionalis (Sergienko 1992) [=Hoplophthiracarus meridionalis] S04
         |--N. (C.) nitidus (Pérez-Íñigo & Baggio 1988) [=Calyptophthiracarus nitidus] S04
         |--N. (C.) oenipontanus (Mahunka 1982) [=Hauserophthiracarus oenipontanus] S04
         |--N. (C.) olivaceus (Jacot 1929) S04 (see below for synonymy)
         |--N. (C.) pavidus (Berlese 1913) S04 (see below for synonymy)
         |    |--N. p. pavidus S04 (see below for synonymy)
         |    `--N. p. inopinatus (Mahunka 1991) [=Calyptophthiracarus cretensis inopinatus] S04
         |--N. (C.) texanus (Niedbała 2002) [=Protophthiracarus texanus] S04
         `--N. (C.) vicinus (Niedbała 1984) [=Hoplophthiracarus vicinus] S04

‘Hoplophorella’ reducta Mahunka 1988 non Steganacarellus reductus Mahunka 1986 (not preoc. if in dif. gen.) [incl. Notophthiracarus (Notophthiracarus) parareductus Mahunka 1992] S04

Notophthiracarus (Calyptophthiracarus) olivaceus (Jacot 1929) S04 [=Phthiracarus olivaceus S04, P. olivaceum N86; incl. P. erinaceus Jacot 1930 S04]

Notophthiracarus (Calyptophthiracarus) pavidus (Berlese 1913) S04 [=Hoploderma pavidum S04, Hoplophthiracarus pavidus N86, Phthiracarus pavidus N86]

Notophthiracarus (Calyptophthiracarus) pavidus pavidus (Berlese 1913) S04 [incl. Hoplophthiracarus cretensis Mahunka 1979 S04, Calyptophthiracarus cretensis N86, Phthiracarus pavidus minor Krivolutsky 1975 N86]

Notophthiracarus (Notophthiracarus) indicus (Bayoumi & Mahunka 1979) S04 [=Hoplophthiracarus indicus S04, Austrophthiracarus indicus N86]

Notophthiracarus (Notophthiracarus) indiligens (Niedbała 1984) S04 [=Hoplophthiracarus indiligens S04, Austrophthiracarus indiligens N86]

Notophthiracarus (Notophthiracarus) inenarrabilis (Niedbała 1982) S04 [=Hoplophthiracarus inenarrabilis S04, Austrophthiracarus inenarrabilis N86]

Notophthiracarus (Notophthiracarus) obsessus Subías 2004 S04 [=N. admirabilis Niedbała & Colloff 1997 non Neophthiracarus admirabilis Niedbała 1982 S04]

Notophthiracarus (Notophthiracarus) rafalskii (Niedbała 1997) S04 [=Arphthicarus rafalskii non N. rafalskii Niedbała 1997 (not preoc. if in dif. gen.) S04, Atropacarus (Hoplophorella) rafalskii N91]

Notophthiracarus (Notophthiracarus) schizocomus (Hammer 1962) S04 [=Steganacarus schizocomus S04, Hoplophthiracarus schizocoma N86; incl. No. niger Mahunka 1980 S04, Neophthiracarus niger N86]

Notophthiracarus (Notophthiracarus) veteratorius (Niedbała 1988) S04 [=Hoplophthiracarus veteratorius S04, Austrophthiracarus veteratorius N86]

*Type species of generic name indicated

REFERENCES

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[N86] Niedbała, W. 1986. Catalogue des Phthiracaroidea (Acari), clef pour la détermination des espèces et descriptions d'espèces nouvelles. Annales Zoologici 40 (4): 309–370.

[N87] Niedbała, W. 1987. Phthiracaroidea (Acari, Oribatida) nouveaux d'Australie. Redia 70: 301–375.

[N91] Niedbała, W. 1991. Preliminary hypotheses on the biogeography of the Phthiracaroidea (Acari, Oribatida). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 219–231. SPB Academic Publishing: The Hague.

[N07] Niedbała, W. 2007. Ptyctimous mites (Acari, Oribatida) of Costa Rica. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 39–42. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 7 March 2019.