New Guinea spiny bandicoot Echymipera kalubu, photographed by Michael Pennay.

Belongs within: Marsupialia.

The Peramelemorphia includes the bandicoots, a group of long-snouted Australian marsupials. Most species are insectivorous or carnivorous, though the now-extinct pig-footed bandicoot Chaeropus ecaudatus may have been a grazer (Long et al. 2002.

Characters (from Long et al. 2002): Lower molars with very large, un-notched anterior cingulid; upper molars with large adjacent stylar cusps C and D, and buccally shifted centrocrista. Hind feet syndactylous (at least in Perameloidea), with second and third toes reduced and joined by a web of skin).

    |--Yarala Muirhead & Filan 1995 [Yaralidae, Yaraloidea] LA02
    |    |--*Y. burchfieldi Muirhead & Filan 1995 LA02
    |    `--Y. kida MJ11
    `--Perameloidea [Peramelidae] LA02
         |--Chaeropus ecaudatus CB-E04
         `--+--Thylacomyidae LA02
            |    |--Ischnodon Stirton 1955 LA02
            |    |    `--*I. australis Stirton 1955 LA02
            |    |--Macrotis CB-E04
            |    |    |--M. lagotis CB-E04
            |    |    `--M. leucura CB-E04
            |    `--Thylacomys nigripes WJ26
            `--+--+--Isoodon CB-E04
               |  |    |--I. macrourus (Gould 1842) CB-E04, K92
               |  |    `--+--I. auratus CB-E04
               |  |       `--I. obesulus (Shaw 1797) CB-E04, BD-D09
               |  `--Perameles Geoffroy 1803 CB-E04, LA02
               |       |  i. s.: P. allinghamensis Archer 1976 LA02
               |       |         P. bowensis Muirhead, Dawson & Archer 1997 LA02
               |       |         P. eremiana CB-E04
               |       |         P. wombeyensis Broom 1896 F71
               |       |--P. bougainville CB-E04
               |       `--+--*P. nasuta Geoffroy 1804 LA02, CB-E04, LA02
               |          `--P. gunnii CB-E04
               `--Peroryctidae LA02
                    |--Rhynchomeles prattorum LA02, BP87
                    |--Peroryctes CB-E04
                    |    |--P. broadbenti CB-E04
                    |    |--P. longicaudata TL70
                    |    `--P. raffrayana CB-E04
                    |--Microperoryctes LA02
                    |    |--M. longicauda CB-E04
                    |    |--M. murina CB-E04
                    |    `--M. papuensis CB-E04
                    `--Echymipera Lesson 1842 LA02
                         |--*E. kalubu (Lesson 1828) [=Perameles kalubu] R64
                         |--E. clara CB-E04
                         |--E. davidi CB-E04
                         |--E. doreyana TL70
                         |--E. echinista CB-E04
                         `--E. rufescens CB-E04
                              |--E. r. rufescens R64
                              `--E. r. australis Tate 1948 R64

*Type species of generic name indicated


[BD-D09] Beveridge, I. & M.-C. Durette-Desset. 2009. A new species of the nematode genus Copemania (Nematoda: Trichostrongylida), parasitic in the western quoll, Dasyurus geoffroii and short-nosed bandicoot, Isoodon obesulus from south-western Western Australia. Records of the Western Australian Museum 25 (3): 345-349.

[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[K92] Klompen, J. S. H. 1992. Phylogenetic relationships in the mite family Sarcoptidae (Acari: Astigmata). Miscellaneous Publications, Museum of Zoology, University of Michigan 180: i-vi, 1-154.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[MJ11] Meredith, R. W., J. E. Janečka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer & W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334: 521-524.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[TL70] Turnbull, W. D. & E. L. Lundelius, Jr. 1970. The Hamilton fauna: a Late Pliocene mammalian fauna from the Grange Burn, Victoria, Australia. Fieldiana Geology 19: 1-163.

[WJ26] Wood Jones, F. 1926. The R. M. Johnston Memorial Lecture, 1925. The mammalian toilet and some considerations arising from it. Papers and Proceedings of the Royal Society of Tasmania 1925: 14-62.

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