Cerambycidae

Callidium violaceum, photographed by Siga.


Belongs within: Phytophaga.
Contains: Stenoderinae, Exocentrinae, Niphoninae, Saperdinae, Erythrinae, Callichrominae, Hypselominae, Leptura, Phytoeciinae, Mesosinae, Apomecyninae, Penthea, Batocera, Rhytiphora, Symphyletes, Praonetha, Glenea, Tmesisternus, Zygocera, Hebecerus, Agapanthiini, Prioninae, Cerambyx, Hesthesis, Phoracantha, Rhinotragini, Uracanthus.

The Cerambycidae include the longicorn beetles, distinguished (as their name suggests) by the notably long antennae of many species, which may be longer than the remainder of the insect. An exception to this, however, are members of the subfamily Parandrinae, in which the antennae rarely extend beyond the posterior margin of the prothorax. Larvae of longicorns are generally internal feeders on plants, and some are significant pests as borers into wood.

Other major subgroups of the longicorns include the Lamiinae, which usually have a strongly deflexed, hypognathous head (Lawrence & Britton 1981). Members of the subfamily Cerambycinae are smaller, relatively long-legged species, often brightly coloured (Zborowski & Storey 2010). The Disteniidae are slender longicorns that have been treated as a separate family owing to differences in their larval morphology but were found nested among the Cerambycidae in the molecular analysis by McKenna et al. (2015).

Characters (from Lawrence & Britton 1991): Elongate, subcylindrical to somewhat flattened beetles, usually pubescent; antennae almost always more than two-thirds body length and inserted on prominences, so that they can be directed backwards over the body. Eyes usually emarginate, often deeply so; lateral pronotal carinae usually absent, often with lateral tubercles or spines on prothorax; mesonotum usually with stridulatory file; tibial spurs present on all legs; tarsal claws simple. Larvae elongate, subcylindrical to slightly flattened, lightly sclerotised and subglabrous or clothed with fine setae or patches of spinules. Head retracted into prothorax and excavated posteriorly, forming recess housing retractor muscle; antennae short; mandibles large and stout; prothorax usually more or less enlarged; legs small and widely separated, may be vestigial or absent; first six or seven abdominal segments may bear fleshy ampullae; urogomphi, if present, minute.

<==Cerambycidae [Cerambicini, Cerambycoidea, Lamiidae, Longicornia, Prionidae]
    |--+--Disteniidae [Disteniinae] MW15
    |  |    |--Cometes hirticornis Lepeletier & Audinet-Serville 1828 B14
    |  |    `--Noemia P66b
    |  |         |--N. chalybeata Pascoe 1866 P66b
    |  |         `--N. flavicornis P66b
    |  `--Lamiinae [Lamiides] B14
    |       |  i. s.: Sarothrocera P66a
    |       |         Diastocera P66a
    |       |         Phryneta caeca P66a
    |       |         Imalmus P66a
    |       |         Hagesata P66a
    |       |         *Leprodera elongata P66a
    |       |         Archidice P66a
    |       |         Euoplia P66a
    |       |         Lamia E61
    |       |           |--L. aedilis L02
    |       |           |--L. araneiformis L02
    |       |           |--L. carcharias [=Saperda carcharias] L02
    |       |           |--L. fuliginator L02
    |       |           |--L. longimana [=Prionus longimanus] L02
    |       |           |--L. sutor L02
    |       |           |--L. textor E61
    |       |           `--L. tristis L02
    |       |         Penthea M87
    |       |         Apriona Chevrolet 1852 P66a
    |       |           `--A. germari [=Lamia germarii] P66a
    |       |         Thestus Pascoe 1866 P66a
    |       |           `--*T. oncideroides Pascoe 1866 P66a
    |       |         Metopides Pascoe 1866 P66a
    |       |           `--*M. occipitalis Pascoe 1866 P66a
    |       |         Epepeotes Pascoe 1866 P66a
    |       |           `--*E. luscus [=Lamia lusca] P66a
    |       |         Blepephaeus Pascoe 1866 P66a
    |       |           `--*B. succinctor (Chevrolat 1852) (see below for synonymy) P66a
    |       |         Epicedia P66a
    |       |           |--*E. pleuricausta [=Leprodera pleuricausta; incl. Lamia carcelii] P66a
    |       |           `--E. plagiata [=Leprodera plagiata; incl. L. trimaculata (n. n.)] P66a
    |       |         Imantocera P66a
    |       |           |--I. penicillata P66a
    |       |           `--I. plumosa [=Cerambyx plumosus] P66a
    |       |         Gnoma P66a
    |       |           |--G. affinis Guér. 1830 M86
    |       |           |--G. dispersa Pascoe 1866 P66a
    |       |           |--G. longicollis P66a [=Lamia longicollis L02]
    |       |           `--G. suturalis Westw. 1832 M87
    |       |         Mecotagus Pascoe 1866 P66a
    |       |           |--*M. tigrinus [=Cerambyx tigrinus] P66a
    |       |           `--M. guerinii [=Pelargoderus guerinii] P66a
    |       |         Batocera P66a
    |       |         Cerosterna [=Celosterna (l. c.)] P66a
    |       |           |--C. approximator P66a
    |       |           |--C. clathrator P66a
    |       |           `--C. reticulator P66a
    |       |         Olenecamptus Chevrolat 1835 P66a
    |       |           |--O. bilobus (see below for synonymy) P66a
    |       |           |--O. optatus Pascoe 1866 [incl. Schoeniocera sexnotata (n. n.)] P66a
    |       |           `--O. quietus Pascoe 1866 P66a
    |       |         Monochamus B14
    |       |           |--M. alternatus Hope 1842 B14
    |       |           |--M. fistulator [=Lamia fistulator] P66a
    |       |           |--M. galloprovincialis [incl. M. galloprovincialis var. cinerascens] C01
    |       |           |--M. musivus Pascoe 1866 P66a
    |       |           |--M. quadrimaculatus C01
    |       |           |--M. sartor C01
    |       |           |--M. sobrius P66a
    |       |           `--M. sutor C01
    |       |         Anoplophora B14
    |       |           |--A. chinensis DAF07
    |       |           |--A. coeruleoantennata B14
    |       |           |--A. freyi B14
    |       |           `--A. glabripennis (Motschulsky 1853) [incl. A. nobilis] B14
    |       |         Doliops B14
    |       |           |--D. curculionoides B14
    |       |           |--D. duodecimpunctata Heller 1923 B14
    |       |           `--D. gutowskii B14
    |       |         Iberodorcadion B14
    |       |           |  i. s.: I. fuliginator (Linnaeus 1758) [=Dorcadion fuliginator] B14
    |       |           |--I. (Baeticodorcadion) mucidum F05
    |       |           |    |--I. m. mucidum F05
    |       |           |    `--I. m. rondensis Verdugo 2003 F05
    |       |           `--I. (Hispanodorcadion) zenete Anichtchenko & Verdugo 2004 F05
    |       |         Zygrita diva Thoms. 1860 [incl. Z. nigrozonata Thoms. 1860] M87
    |       |         Microtragus LB91
    |       |           |--M. arachne M87
    |       |           |--M. eremita M87
    |       |           |--M. luctuosus Schuck. 1838 M87
    |       |           |--M. mormon M87
    |       |           |--M. senex White 1846 M87
    |       |           |--M. sticticus Pascoe 1866 M87
    |       |           `--M. waterhousei Pascoe 1863 M87
    |       |         Athemistus LB91
    |       |           |--A. aethiops Pascoe 1867 M87
    |       |           |--A. armitagei Pascoe 1866 M87
    |       |           |--A. bituberculatus Pascoe 1867 M87
    |       |           |--A. funereus Pascoe 1866 M87
    |       |           |--A. howittii Pascoe 1867 M87
    |       |           |--A. pubescens Pascoe 1862 M87
    |       |           |--A. puncticollis Pascoe 1867 M87
    |       |           `--A. rugosulus Guér. 1830 [incl. A. tuberculatus] M87
    |       |         Rosenbergia megacephala v. d. Poll 1886 M87
    |       |         Ancita crossotoides Thoms. 1864 M87
    |       |         Platyomopsis LB91
    |       |           |--P. armatula White 1859 M87
    |       |           |--P. obliqua Donov. 1805 [incl. P. spinosa] M87
    |       |           `--P. tuberculata Hope 1841 M87
    |       |         Rhytiphora C91
    |       |         Pithomictus decoratus M86
    |       |         Aegomomus M86
    |       |           |--A. insularis [incl. Coptops abdominalis] P66a
    |       |           `--A. ornatus Macleay 1886 M86
    |       |         Escharodes paganus M86
    |       |         Symphyletes M87
    |       |         Sotades platypus M86
    |       |         Praonetha M86
    |       |         Dihammus longicornis M86
    |       |         Glenea M86
    |       |         Oberea Mulsant 1839 M86, P66a [incl. Isosceles Newman 1840 P66a]
    |       |           |--O. clara Pascoe 1866 P66a
    |       |           |--O. curialis Pascoe 1866 P66a
    |       |           |--O. mundula M86
    |       |           |--O. pigra Newm. 1851 M87
    |       |           `--O. tenuata Pascoe 1866 P66a
    |       |         Hestima trigeminata M86
    |       |         Orinoeme rubricollis Macleay 1886 M86
    |       |         Elais bimaculata M86
    |       |         Sphingnotus M87
    |       |           |--S. insignis Perroud 1855 M87
    |       |           `--S. mirabilis [=Tmesisternus mirabilis] M86
    |       |         Tmesisternus M86
    |       |         Temnosternus M87
    |       |           |--T. apicalis Pascoe 1878 M87
    |       |           |--T. dissimilis Pascoe 1859 M87
    |       |           |--T. planiusculus M87
    |       |           `--T. vitulus Pascoe 1871 M87
    |       |         Dorcadida biocularis White 1846 M87
    |       |         Stychus amycteroides Pascoe 1859 [incl. S. pascoei] M87
    |       |         Ceraegidion horrens M87
    |       |         Corestetha insularis Pascoe 1875 M87
    |       |         Mesolita M87
    |       |           |--M. lineolata M87
    |       |           `--M. transversa M87
    |       |         Monohammus M87
    |       |           |--M. acanthias Pascoe 1875 M87
    |       |           |--M. desperatus M87
    |       |           |--M. fasciatus Montrouz. 1855 [incl. M. argutus] M87
    |       |           |--M. fistulator Germ. 1824 M87
    |       |           |--M. mixtus Hope 1841 M87
    |       |           |--M. ovinus Pascoe 1863 M87
    |       |           |--M. togatus Perroud 1855 M87
    |       |           `--M. vastator Newm. 1847 M87
    |       |         Pelargoderus arouensis Thoms. 1857 M87
    |       |         Callipyrga turrita [incl. C. pallidicornis] M87
    |       |         Meton M87
    |       |           |--M. digglesi Pascoe 1859 M87
    |       |           `--M. fasciatus Pascoe 1878 M87
    |       |         Arsysia vittata Gestro 1876 M87
    |       |         Dysthaeta anomala Pascoe 1859 M87
    |       |         Oricopis umbrosus Pascoe 1863 M87
    |       |         Demonassa macleayi Pascoe 1859 [incl. D. funeraria Thoms. 1864] M87
    |       |         Zygocera M87
    |       |         Cyocyphax praonetoides Thoms. 1878 M87
    |       |         Thyada barbicornis Pascoe 1859 M87
    |       |         Probatodes piliger Macleay 1827 [incl. P. plumula Newm. 1851] M87
    |       |         Velora sordida Pascoe 1863 [incl. V. australis Thoms. 1864] M87
    |       |         Hebecerus M87
    |       |         Lychrosis M87
    |       |           |--L. afflictus Pascoe 1867 M87
    |       |           `--L. luctuosus Pascoe 1863 M87
    |       |         Hathliodes M87
    |       |           |--H. costulatus Pascoe 1867 M87
    |       |           |--H. grammicus Pascoe 1859 M87
    |       |           |--H. lacteolus (Hope 1841) [=Hathlia lacteola] M87
    |       |           |--H. lineellus (Hope 1841) [=Hathlia lineella] M87
    |       |           |--H. melanocephalus (Hope 1841) (see below for synonymy) M87
    |       |           |--H. moratus Pascoe 1866 M87
    |       |           |--H. murinus Pascoe 1859 M87
    |       |           `--H. quadrilineatus (Hope 1841) [=Hathlia quadrilineata] M87
    |       |         Micracantha M87
    |       |           |--M. abdominalis M87
    |       |           |--M. irata (Pascoe 1862) [=Aegomomus iratus] M87
    |       |           |--M. misella (Pascoe 1863) [=Aegomomus misellus] M87
    |       |           |--M. oblita (Pascoe 1863) [=Aegomomus oblitus] M87
    |       |           |--M. torosa (Pascoe 1863) [=Aegomomus torosus] M87
    |       |           `--M. woodlarkiana Montrouz. 1855 [incl. M. bakewelli Pascoe 1859, M. porosa] M87
    |       |         Chaetostigme casta Pascoe 1875 M87
    |       |         Prosoplus hollandicus Boisd. 1835 M87
    |       |         Atyporus intercalaris Pascoe 1867 M87
    |       |         Menyllus maculicornis Pascoe 1864 [incl. Sysspilotus macleayi Pascoe 1865] M87
    |       |         Acriotypa basalis Pascoe 1875 M87
    |       |         Saperdopsis armata Thoms. 1864 M87
    |       |         Depsages granulosa Guér. 1830 M87
    |       |         Iphiastus heros Pascoe 1863 M87
    |       |         Corrhenes M87
    |       |           |--C. cruciata Pascoe 1875 M87
    |       |           |--C. fulva Pascoe 1875 M87
    |       |           |--C. funesta Pascoe 1859 M87
    |       |           |--C. grisella Pascoe 1875 M87
    |       |           |--C. guttulata M87
    |       |           |--C. mystica Pascoe 1863 M87
    |       |           |--C. paulla Germ. 1848 M87
    |       |           `--C. stigmatica M87
    |       |         Mycerinopsis M87
    |       |           |--M. arida (Pascoe 1862) [=Mycerinus aridus] M87
    |       |           `--M. uniformis (Pascoe 1863) [=Mycerinus uniformis] M87
    |       |         Eunidia australica Thoms. 1868 M87
    |       |         Anaesthetis lepida Germ. 1848 M87
    |       |         Phaeapete M87
    |       |           |--P. albula Pascoe 1865 M87
    |       |           `--P. denticollis Pascoe 1867 M87
    |       |         Bucynthia spiloptera Pascoe 1863 M87
    |       |         Essisus dispar Pascoe 1866 M87
    |       |         Itheum M87
    |       |           |--I. lineare Pascoe 1864 M87
    |       |           `--I. vittigerum Pascoe 1864 M87
    |       |         Lagochirus incei Newm. 1847 M87
    |       |         Neissa M87
    |       |           |--N. inconspicua Pascoe 1866 M87
    |       |           `--N. nigrina Pascoe 1866 M87
    |       |         Pentacosmia scoparia Newm. 1842 M87
    |       |         Illaena exilis Erichs. 1842 M87
    |       |         Ameipsis marginicollis Pascoe 1865 M87
    |       |--Acrocinus longimanus (Linnaeus 1758) B14
    |       |--Agapanthiini B14
    |       |--Cyrtinus [Cyrtinini] B14
    |       |    `--C. pygmaeus (Haldeman 1847) B14
    |       |--Oncideres [Onciderini] B14
    |       |    `--O. cingulata (Say 1826) B14
    |       |--Onychocerus [Anisocerini] B14
    |       |    `--O. albitarsis Pascoe 1859 B14
    |       |--Petrognatha [Petrognathini] B14
    |       |    `--P. gigas Fabricius 1792 B14
    |       |--Acanthoderini B14
    |       |    |--Macropophora B14
    |       |    `--Acanthoderes jaspidea [incl. A. albifrons Sturm. 1826, A. costata] M87
    |       |--Moneilema [Moneilemini] B14
    |       |    |--M. annulatum Say 1824 B14
    |       |    `--M. ulkei G25
    |       |--Tetraopes MW15 [Tetraopini B14]
    |       |    |--T. femoratus LeConte 1847 B14
    |       |    `--T. tetrophthalmus M73
    |       `--Ceroplesini B14
    |            |--Ceroplesis elegans Gestro 1889 G89
    |            `--Paranaleptes Breuning 1937 B14, O06
    |                 `--*P. reticulata (Thomson 1877) O06
    `--+--+--+--Prioninae MW15
       |  |  `--Parandrinae B14
       |  |       |--Parandra Latreille 1802 MW15, L02
       |  |       |    |--*P. glabra [=Attelabus glaber] L02
       |  |       |    `--P. frenchi L19
       |  |       `--Neandra B14
       |  |            |--N. brunnea (Fabricius 1798) B14
       |  |            `--N. marginicollis B14
       |  `--+--Rutpela MW15
       |     `--+--Vesperus [Vesperidae, Vesperinae] B14
       |        |    |--V. gomezi Verdugo 2004 F05
       |        |    `--V. luridus (Rossi 1794) B14
       |        `--Dorcasominae B14
       |             |--Apatophysis [Apatophyseinae] MW15
       |             |    `--A. serricornis (Gebler 1843) [=Pachyta serricornis; incl. A. mongolica] B14
       |             `--Capnolymma B14
       |                  |--*C. stygia Pascoe 1858 B14
       |                  `--C. capreola Pascoe 1866 P66b
       `--Cerambycinae [Cerambycides, Clytinae] SW03
            |  i. s.: Dialeges pauper P66b
            |         Pachydissus P66b
            |           |--P. nubilus Pascoe 1863 M87
            |           |--P. picipennis Germ. 1848 M87
            |           `--P. sericus [incl. P. turbinaticornis] M87
            |         Conothorax P66b
            |         Tapinolachnus P66b
            |         Calpazia P66b
            |         Lachnopterus P66b
            |         Aprosictus Pascoe 1866 P66b
            |         Ambeodontus tristis M59
            |         Aphiorrhynchus M87
            |           |--A. apicalis Pascoe 1859 M87
            |           |--A. divisus Pascoe 1871 M87
            |           `--A. pulcher Hope 1834 [incl. Psilomorpha lusoria Pascoe 1865] M87
            |         Aridaeus M87
            |           |--A. heros Pascoe 1867 M87
            |           `--A. thoracicus Don. 1805 M87
            |         Cyriopalus Pascoe 1866 P66b
            |           `--*C. wallacei Pascoe 1866 P66b
            |         Hoplocerambyx P66b
            |           |--H. morosus P66b
            |           |--H. relictus Pascoe 1866 P66b
            |           `--H. spinicornis Z94
            |         Rhytidodera P66b
            |           |--R. cristata Pascoe 1866 P66b
            |           `--R. simulans [=Hammaticherus simulans] P66b
            |         Cerambyx P66b
            |         Neocerambyx P66b
            |           |--*N. paris P66b
            |           |--N. aurifaber P66b
            |           |--N. intricatus Pascoe 1866 P66b
            |           `--N. lambii Pascoe 1866 P66b
            |         Imbrius Pascoe 1866 P66b
            |           |--I. ephebus Pascoe 1866 P66b
            |           |--I. lineatus Pascoe 1866 P66b
            |           |--I. micaceus [=Cerambyx micaceus] P66b
            |           `--I. strigosus Pascoe 1866 P66b
            |         Coptomma Newman 1840 (see below for synonymy) SW03
            |           |  i. s.: ‘Navomorpha’ douei Lucas 1863 SW03
            |           |--C. variegatum (Fabricius 1775) (see below for synonymy) SW03
            |           `--+--C. sulcatum (Fabricius 1775) (see below for synonymy) SW03
            |              `--+--C. marrisi Song & Wang 2003 SW03
            |                 `--+--C. lineatum (Fabricius 1775) (see below for synonymy) SW03
            |                    `--C. sticticum (Broun 1893) [=Navomorpha sticticum, N. stictica (l. c.)] SW03
            |         Phymatodes B14
            |           |--P. decussatus B14
            |           `--P. nitidus LeConte 1874 B14
            |         Rosalia alpina (Linnaeus 1758) B14
            |         Epithora LB91
            |           |--E. angustata Boisd. 1835 M87
            |           |--E. dorsalis [incl. E. elongata, E. rhombifera, E. uniguttata] M87
            |           `--E. undulata Hope 1843 [incl. E. rubripes Hope 1843] M87
            |         Bardistus cibarius Newm. 1841 [incl. Dicranops piceus] M87
            |         Hesthesis LB91
            |         Macrones LB91
            |           |--M. acicularis Pascoe 1862 M87
            |           |--M. capito Pascoe 1863 M87
            |           |--M. exilis M87
            |           |--M. rufus M87
            |           `--M. subclavatus Pascoe 1871 M87
            |         Eroschema LB91
            |           |--E. atricolle Pascoe 1865 M87
            |           `--E. poweri Pascoe 1859 [incl. Octavia nigricolle] M87
            |         Diotimana undulata LB91
            |         Ceresium [Eburiinae] M86
            |           |--C. annulicorne Germ. 1848 M87
            |           |--C. pachymerum M86
            |           |--C. raripilum P66b
            |           |--C. simplex P66b (see below for synonymy)
            |           |--C. versutum Pascoe 1866 P66b
            |           |--C. vestigiale Pascoe 1866 P66b
            |           `--C. zeylanicum P66b
            |         Tethionea strumosa M86
            |         Chloridolum M86
            |           |--C. cinderellum M87
            |           |--C. cinnyris Pascoe 1866 P66b
            |           |--C. dorycum M86
            |           `--C. thomsoni [=Callichroma thomsoni] P66b
            |         Demonax macilenta [=Acrocyrta macilenta] P66b
            |         Epipedocera P66b
            |         Homalomelas P66b
            |         Prothema P66b
            |         Obrida P66b
            |           |--O. comata Pascoe 1863 M87
            |           `--O. fascialis White 1846 M87
            |         Typhocesis macleayi Pascoe 1863 M87
            |         Clytus M86
            |           |--C. curtisi M87
            |           |--C. durvillei M87
            |           |--C. sexguttatus Lucas 1847 E12
            |           |--C. sexmaculatus Donov. 1805 M87
            |           `--C. velutinus Macleay 1886 M86
            |         Bicon Pascoe 1866 P66b
            |           `--*B. sanguineus Pascoe 1866 P66b
            |         Sigeum Pascoe 1866 P66b
            |           `--*S. humerale [=Blemmya humeralis] P66b
            |         Asmedia Pascoe 1866 P66b
            |           `--*A. mimetes Pascoe 1866 P66b
            |         Dere P66b
            |           |--*D. thoracica P66b
            |           `--D. marginata Pascoe 1866 P66b
            |         Xylotrechus Chevrolat 1860 P66b
            |           |--X. australis P66b [=Clytus australis P66b; incl. C. phidias Newm. 1842 M87]
            |           |--X. ibex C01
            |           `--X. rusticus C01
            |         Clytanthus [incl. Anthoboscus, Chlorophorus] P66b
            |           |--C. angustulus Macleay 1886 M86
            |           |--C. annularis [=Callidium annulare, *Chlorophorus annularis] P66b
            |           |--C. glaucinus [=Perissus glaucinus] P66b
            |           |--C. latofasciatus C01
            |           `--C. sumatrensis P66b
            |         Euryarthrum Blanchard 1845 [incl. Blemmya Pascoe 1856] P66b
            |           |--E. albocinctum P66b
            |           |--E. atripenne Pascoe 1866 P66b
            |           |--E. bifasciatum P66b
            |           |--E. carinatum Pascoe 1866 P66b
            |           |--E. egenum Pascoe 1866 P66b
            |           |--E. interruptum Pascoe 1866 P66b
            |           |--E. lambii Pascoe 1866 P66b
            |           `--E. nodicolle Pascoe 1866 P66b
            |         Hammatochaerus australasiae Hope 1842 M87
            |         Saporaea femoralis Thoms. 1878 M87
            |         Phacodes M87
            |           |--P. distinctus Pascoe 1865 M87
            |           |--P. elusus Pascoe 1865 M87
            |           |--P. essingtoni Hope 1841 M87
            |           |--P. ferrugineus Pascoe 1863 M87
            |           |--P. fuscus Pascoe 1865 M87
            |           |--P. longicollis Pascoe 1871 M87
            |           |--P. mossmanni Newm. 1851 M87
            |           |--P. obscurus [incl. P. lentiginosus] M87
            |           `--P. tenuitarsis Pascoe 1871 M87
            |         Oebarina M87
            |           |--O. ceresioides Pascoe 1866 M87
            |           `--O. tristis Pascoe 1866 M87
            |         Aesiotyche favosa M87
            |         Hypermallus atomarius Drury 1773 [incl. Thersalus bispinosus Pascoe 1863] M87
            |         Atesta M87
            |           |--A. angasi Pascoe 1863 M87
            |           |--A. balteata Pascoe 1863 M87
            |           `--A. bifasciata Pascoe 1863 M87
            |         Allotisis M87
            |           |--A. discreta Pascoe 1863 M87
            |           |--A. scitula Pascoe 1863 M87
            |           `--A. unifasciata Hope 1843 M87
            |         Coptocercus M87
            |           |--C. aberrans Newm. 1841 M87
            |           |--C. biguttatus Donov. 1805 (see below for synonymy) M87
            |           |--C. decorus Perroud 1855 M87
            |           |--C. pedator Pascoe 1863 M87
            |           |--C. politus Pascoe 1863 M87
            |           |--C. pubescens Pascoe 1863 M87
            |           `--C. rubripes Boisd. 1835 (see below for synonymy) M87
            |         Demelius semirugosus Waterh. 1874 M87
            |         Thoris eburifera Pascoe 1867 M87
            |         Ectinope spinicollis Pascoe 1875 M87
            |         Sisyrium M87
            |           |--S. dorsale (Pascoe 1863) [=Obrium dorsale] M87
            |           |--S. ibidionoides (Pascoe 1859) [=Obrium ibidionoides] M87
            |           |--S. stigmosum [=Igenia stigmosa] M87
            |           `--S. tripartitum [=Obrium tripartitum] M87
            |         Psylacrida gracilis Thoms. 1878 M87
            |         Cleistimum venatum Thoms. 1878 M87
            |         Acyrusa ciliata Pascoe 1863 M87
            |         Skeletodes tetrops Newm. 1850 [incl. Elete inscriptus Pascoe 1863] M87
            |         Sidis opiloides M87
            |         Paphora modesta Pascoe 1863 M87
            |         Porithea intorta Newm. 1842 M87
            |         Bethelium M87
            |           |--B. inscriptum Pascoe 1862 M87
            |           `--B. signiferum Newm. 1840 (see below for synonymy) M87
            |         Adrium M87
            |           |--A. artifex Newm. 1842 [incl. A. cucujus, A. terebrans Newm. 1842] M87
            |           `--A. catoxanthum M87
            |         Thephantes clavatus Pascoe 1867 M87
            |         Neostenus M87
            |           |--N. morio (Pascoe 1865) [=Lysestia morio] M87
            |           `--N. saundersii [incl. Lysestia rotundicollis] M87
            |         Aposites M87
            |           |--A. macilentus M87
            |           `--A. pubicollis Pascoe 1866 M87
            |         Mystrosa rubiginea M87
            |         Aphanasium australe Boisd. 1835 [incl. Solimnia sublineatum Pascoe 1863] M87
            |         Tricheops ephippigera M87
            |         Tessaromma M87
            |           |--T. sericans Erichs. 1842 M87
            |           |--T. triste [incl. T. moestum] M87
            |           `--T. undatum [incl. T. macleayi Hope 1840] M87
            |         Petalodes M87
            |           |--P. laminosus Newm. 1840 M87
            |           `--P. plagiatus White 1858 M87
            |         Opsidota M87
            |           |--O. albipilosa M87
            |           `--O. infecta M87
            |         Piesarthrius M87
            |           |--P. laminosus MC13
            |           `--P. marginellus Hope 1840 M87
            |         Strongylurus M87
            |           |--S. ceresioides Pascoe 1867 M87
            |           |--S. cretifer Hope 1835 [incl. Isalium odewahni] M87
            |           |--S. orbatus M87
            |           |--S. scutellatus (Hope 1840) [=Coptopterus scutellatus] M87
            |           `--S. thoracicus Pascoe 1857 M87
            |         Exaereta unicolor M87
            |         Lygesis M87
            |           |--L. cylindricollis Pascoe 1859 M87
            |           `--L. medica Pascoe 1875 M87
            |         Bebius filiformis Pascoe 1865 M87
            |         Oxymagis grayi Pascoe 1866 M87
            |         Scolecobrotus westwoodi Hope 1835 M87
            |         Emenica nigripennis Pascoe 1875 M87
            |         Aethiora fuliginea Pascoe 1863 M87
            |         Stephanops M87
            |           |--S. marginipennis Fairm. 1879 M87
            |           |--S. nasuta [incl. S. elongaticeps Blanch. 1853] M87
            |           `--S. striicollis Fairm. 1879 M87
            |         Psilomorpha tenuipes Saund. 1850 M87
            |         Tritocosmia M87
            |           |--T. atricilla Newm. 1850 M87
            |           |--T. digglesi Pascoe 1859 M87
            |           |--T. latecostata Fairm. 1879 M87
            |           |--T. paradoxa Pascoe 1859 M87
            |           |--T. roei Hope 1836 M87
            |           `--T. rubea M87
            |         Tropocalymma dimidiatum (Newm. 1841) [=Tropis dimidiata] M87
            |         Tropis oculifera Newm. 1840 M87
            |         Syllitus M87
            |           |--S. bipunctatus Waterh. 1877 M87
            |           |--S. deustus Newm. 1841 M87
            |           |--S. grammicus Newm. 1840 [incl. S. rectus Newm. 1841] M87
            |           |--S. parryi Pascoe 1862 M87
            |           |--S. tabidus Pascoe 1871 M87
            |           `--S. terminatus Pascoe 1871 M87
            |         Demomisis filum Pascoe 1867 M87
            |         Simocrysa discolor Pascoe 1871 M87
            |         Enchoptera M87
            |           |--E. apicalis M87
            |           `--E. nigricornis M87
            |         Oroderes humeralis Saund. 1850 M87
            |         Brachopsis M87
            |           |--B. concolor Saund. 1850 M87
            |           `--B. nupera Pascoe 1863 M87
            |         Pseudocephalus M87
            |           |--P. arietinus Newm. 1851 M87
            |           |--P. formicides Newm. 1842 M87
            |           `--P. mirus Pascoe 1865 M87
            |         Aphneope sericata Pascoe 1863 M87
            |         Zoedia M87
            |           |--Z. divisa Pascoe 1862 M87
            |           |--Z. triangularis Pascoe 1862 M87
            |           `--Z. v-album (Boisd. 1835) [=Clytus v-album; incl. Z. elegans Waterhouse 1878] M87
            |         Bimia M87
            |           |--B. bicolor White 1850 M87
            |           `--B. femoralis M87
            |         Aciptera M87
            |           |--A. semiflava Saund. 1850 M87
            |           `--A. waterhousei Pascoe 1863 M87
            |         Agapete M87
            |           |--A. carissima Newm. 1845 M87
            |           |--A. kreusleri Pascoe 1863 M87
            |           `--A. vestita Pascoe 1866 M87
            |         Molorchus M87
            |           |--M. abbreviatus L02
            |           `--M. sidus (Newm. 1840) [=Heliomanes sidus] M87
            |         Earinus M87
            |           |--E. kreuslerae Pascoe 1866 M87
            |           |--E. mimula Pascoe 1863 M87
            |           `--E. picta Pascoe 1871 M87
            |         Mecynopus M87
            |           |--M. cothurnatus Erichs. 1842 M87
            |           `--M. semivitreus Pascoe 1859 M87
            |         Omotes M87
            |           |--O. cucujides Newm. 1842 M87
            |           |--O. erosicollis Pascoe 1859 M87
            |           `--O. punctatissima Newm. 1851 M87
            |         Phalota M87
            |           |--P. collaris Pascoe 1866 M87
            |           `--P. tenella Pascoe 1863 M87
            |         Nenenia aurulenta Pascoe 1866 M87
            |         Xystoena vittata Pascoe 1866 M87
            |         Trichomesia newmani [incl. Chaeropsis dimidiata] M87
            |         Anteros M87
            |         Distichocera M87
            |           |--D. kirbyi Newm. 1851 M87
            |           |--D. macleayi Newm. 1851 M87
            |           |--D. maculicollis [incl. D. ferruginea, D. fulvipennis, D. rubripennis Macleay 1827] M87
            |           |--D. mutator M87
            |           |--D. par Newm. 1851 [incl. D. fuliginosa] M87
            |           `--D. thomsonella White 1859 M87
            |         Chaodalis macleayi Pascoe 1865 M87
            |         Pytheus M87
            |           |--P. jugosus Newm. 1840 M87
            |           |--P. latebrosus Newm. 1840 [incl. P. pallidus] M87
            |           `--P. pulcherrimus Pascoe 1859 M87
            |         Brachytria M87
            |           |--B. centralis Pascoe 1863 M87
            |           |--B. gulosa Newm. 1840 [incl. Callidiomorphus depressa] M87
            |           |--B. picta Waterhouse 1877 M87
            |           `--B. varia Waterhouse 1877 M87
            |         Pempsamacra M87
            |           |--P. dispersa Newm. 1842 M87
            |           |--P. pygmaea Newm. 1851 [incl. Lepidisia bimaculata White 1855] M87
            |           |--P. subaurea Pascoe 1863 M87
            |           |--P. tillides Newm. 1838 M87
            |           `--P. vestita Pascoe 1859 M87
            |         Titurius calcaratus Pascoe 1875 M87
            |         Omophaena M87
            |           |--O. kreusleri Pascoe 1863 M87
            |           `--O. taeniata Pascoe 1866 M87
            |         Schizopleurus balteatus Lacord. 1869 M87
            |         Telocera wollastoni White 1858 M87
            |         Eburigera octoguttata White 1855 M87
            |         Taphos aterrimus Pascoe 1863 M87
            |         Bixorestes M87
            |           |--B. dectus White 1855 M87
            |           `--B. interruptus Oliv. 1790 M87
            |         Hemisthocera flavilinea Newm. 1850 M87
            |         Cremys diophthalmus Pascoe 1862 M87
            |         Ochyra coarctata Pascoe 1871 M87
            |         Homaemota M87
            |           |--H. basalis Pascoe 1865 M87
            |           `--H. duboulayi M87
            |         Ectosticta M87
            |           |--E. cleroides White 1855 [incl. E. simillima White 1855] M87
            |           |--E. eburata Pascoe 1863 M87
            |           `--E. ruida Pascoe 1866 M87
            |         Ipomoria tillides Pascoe 1866 M87
            |         Sophron M87
            |           |--S. eburatus Pascoe 1865 M87
            |           `--S. inornatus Newm. 1842 [incl. Cleadne melanaria Pascoe 1863] M87
            |         Amphirhoe decora Newm. 1841 M87
            |         Leptocera M87
            |           |--L. suturalis (Pascoe 1867) [=Glaucytes suturalis] M87
            |           `--L. vittifera M87
            |         Acrocyrta chrysoderes (White 1855) [=Clytanthus chrysoderes] M87
            |         Ischnotes M87
            |           |--I. bakewelli Pascoe 1859 M87
            |           `--I. cylindracea Newm. 1840 M87
            |         Tragocerus M87
            |           |--T. bidentatus Donov. 1805 [incl. T. australis Serv. 1834] M87
            |           |--T. fasciatus Donov. 1805 [incl. T. bifasciatus Guér. 1830] M87
            |           |--T. formosus Pascoe 1862 M87
            |           |--T. lepidopterus Schreib. 1802 M87
            |           |--T. spencei Hope 1834 M87
            |           `--T. subfasciatus Germ. 1848 M87
            |         Anastetha raripila Pascoe 1866 M87
            |         Sternaderus B70
            |         Pterostenus B70
            |--Phoracantha MW15
            |--Rhinotragini B14
            |--Uracanthus TW07
            |--Hylotrupes [Hylotrupini] B14
            |    |--H. asemoides Pascoe 1863 M87
            |    `--H. bajulus (Linnaeus 1758) B14
            |--Rhopalophorini B14
            |    |--Rhopalophora P66b
            |    `--Cosmisoma ammiralis (Linnaeus 1767) B14
            |--Rhagiomorpha M87 [Rhagiomorphinae, Rhagiomorphini SW03]
            |    |--R. concolor Macleay 1827 [incl. R. lepturoides, Physodroma sexcostata, R. sordida] M87
            |    |--R. exilis Pascoe 1859 M87
            |    |--R. plagiata Hope 1842 M87
            |    `--R. unicolor Hope 1842 M87
            |--Phlyctaenodes M87 [Phlyctaenodini SW03]
            |    |--P. binodosus Lacord. 1869 M87
            |    |--P. pilosus Pascoe 1863 M87
            |    |--P. pustulatus (Hope 1840) [=Trachelorachys pustulatus] M87
            |    |--P. pustulosus [incl. P. brunneus, P. fumicolor Hope 1840, P. pubescens] M87
            |    `--P. tristis M87
            |--Tillomorphini B14
            |    |--Tetranodus B14
            |    |--Pentanodes B14
            |    |--Tillomorpha moestula M87
            |    `--Euderces reichei LeConte 1873 B14
            |         |--E. r. reichei B14
            |         `--E. r. exilis B14
            |--Callidium C01 [Callidiini B14]
            |    |--C. arietis [=Clytus arietis] L02
            |    |--C. bajulus L02
            |    |--C. erosum Macleay 1855 M87
            |    |--C. funestum Boisd. 1835 M87
            |    |--C. morosum Boisd. 1835 M87
            |    |--C. vexatum Newm. 1847 M87
            |    |--C. violaceum C01
            |    `--C. vittigerum Newm. 1847 M87
            `--Callidiopini SW03
                 |--Didymocantha SW03
                 |    |--D. brevicollis M87
                 |    |--D. obliqua Newm. 1840 [incl. D. varicornis Hope 1840] M87
                 |    `--D. picta SW03
                 `--Callidiopis M87
                      |--C. mutica Lacord. 1869 M87
                      |--C. praecox Erichs. 1842 M87
                      `--C. scutellaris M87

Cerambycidae incertae sedis:
  Ochrocydus huttoni FS01
  Hybolasius vegetus Broun 1881 E02
  Xylotoloides trigonellaris E02
  Ptinosoma E02
  Calliprason pallidum (Pascoe 1875) E02
  Somatidia WH02
  Notorrhina Z02
  Oemona hirta (Fabricius 1775) H82
  Psilocnaeia WFS04
    |--P. nana WFS04
    `--P. parvula WFS04
  Xyloteles WFS04
    |--X. griseus WFS04
    `--X. humeratus WFS04
  Zorion minutum T27
  Neocalliprason Brookes 1927 B27
    `--*N. elegans Brookes 1927 B27
  Pseudocalliprason B27
  Cerambycomima GE05
  Tithoes G89
    |--T. arabicus Thoms. 1877 G89
    `--T. capensis G89
  Obriaccum G89
    |--O. elegans G89
    `--O. manzonii Gestro 1889 G89
  Dichostates G89
    |--D. strigifrons Fairm. 1885 G89
    |--D. subocellatus G89
    `--D. vagepictus G89
  Ceratites jaspideus Serv. 1835 G89
  Rhodopina lewisii IK05
  Spintheriini SW03
  Glaucytes [Glaucytini] SW03
    `--G. interrupta SW03
  Gastrosarus B88
  Poecilipe medialis B88
  Plagithmysus O94
  Mecas C90
  Crossidius C90
  Rhagium mordax C01
  Pachyta quadrimaculata C01
  Brachyta interrogationis [incl. B. interrogationis var. punctata] C01
  Acmaeops C01
    |--A. marginata C01
    `--A. pratensis C01
  Gaurotes virginea [incl. G. virginea var. thalassina] C01
  Oedecnema dubia C01
  Allosterna tabacicolor C01
  Cyrtoclytus C01
    |--C. capra C01
    `--C. monticallisus Komiya 1980 Iw92
  Neodorcadion C01
    |--N. involvens C01
    |--N. orygi C01
    |--N. potanini C01
    `--N. zichyi Csiki 1901 C01
  Aderpas P66a
  Anamera P66a
  Liopus C01
    |--L. femoratus C01
    `--L. ganglbaueri Csiki 1901 C01
  Pogonochaerus fasciculatus C01
  Haplocnemia myops C01
  Erioderus P66b
  Tragosoma P66b
  Paraglenea Bates 1866 B66
    |--*P. fortunei [=Glenea fortunei] B66
    `--P. swinhoei Bates 1866 B66
  Adesmus druryi Im92
  Semanotus Mulsant 1839 CW92
  Atossa [Onocephalinae] P66a
    |--*A. strenua P66a
    `--A. atomaria Pascoe 1866 P66a
  Obriinae P66b
    |--Deuteromma testaceum P66b
    `--Ciopera Pascoe 1866 P66b
         `--*C. decolorata Pascoe 1866 P66b
  Necydalinae B14
    |--Ulochaetes leoninus LeConte 1854 B14
    |--Merionoeda P66b
    |    |--M. acuta Pascoe 1866 P66b
    |    `--M. scitella P66b
    `--Necydalis Linnaeus 1758 L58
         |--N. auricoma Newm. 1840 M87
         |--N. major Linnaeus 1758 L58
         |--N. minor Linnaeus 1758 L58
         `--N. rufa L02
  Astathes Newman 1842 [incl. Tetraophthalmus Blanchard 1845; Astatheinae] P66a
    |--A. nigricornis P66a
    |--A. perplexa P66a
    |--A. splendida [=Cerambyx splendidus] P66a
    `--A. terminata P66a
  Purpuriceninae P66b
    |--Euryclea cardinalis [=Eurycephalus cardinalis] P66b
    |--Euryphagus maxillosus [=Cerambyx maxillosus; incl. C. nigripes] P66b
    `--Purpuricenus [incl. Cyclodera] P66b
         |--P. angasi (White 1847) [=Cyclodera angasi] M87
         |--P. dumerilii Lucas 1847 E12
         |--P. quadrinotatus White 1846 [incl. P. australicus] M87
         `--P. sanguinolentus [=Cerambyx sanguinolentus] P66b
  Acanthocininae P66a
    |--Eoporis elegans P66a
    |--Ostedes P66a
    `--Acanthocinus Z02
         |--A. aedilis C01
         |--A. griseus C01
         |--A. lineola Newm. 1851 M87
         `--A. plumula Newm. 1851 M87
  Spondylidinae [Aseminae] B14
    |--Spondylis buprestoides (Linnaeus 1758) B14
    |--Neospondylis upiformis [=Spondylis upiformis] B14
    |--Arhopalus syriacus LB91
    |--Asemum striatum B14, C01
    `--Tetropium castaneum B14, C01
  Cerasphorinae P66b
    |--Stromatium asperulum P66b
    |--Noserius tibialis P66b
    |--Gnatholea eburifera P66b
    `--Xystrocera P66b
         |--X. alcyonea Pascoe 1866 P66b
         |--X. australasiae Hope 1841 M87
         |--X. globosa [=Cerambyx globosus] P66b
         `--X. virescens Newm. 1840 M87
  Stenoderinae P66b
  Aegosominae P66b
    |--Aegosoma P66b
    |    |--A. marginale [=Cerambyx marginalis] P66b
    |    `--A. scabricorne P66b
    `--Megopis P66b
         |--*M. mutica P66b
         |--M. costipennis P66b
         `--M. procera Pascoe 1866 P66b
  Dorcadioninae P66a
    |--Obages Pascoe 1866 P66a
    |    `--*O. palparis Pascoe 1866 P66a
    `--Dorcadion C01
         |--D. carinatum C01
         |--D. elegans C01
         |--D. equestre C01
         |--D. ribbei [incl. D. ribbei var. rubrofemoratum] C01
         |--D. scabricolle C01
         `--D. sericatum [incl. D. sericatum var. micans] C01
  Exocentrinae P66a
  Niphoninae P66a
  Saperdinae P66a
  Erythrinae P66b
  Callichrominae P66b
  Hypselominae P66a
  Lepturinae P66b
    |--Leptura E12
    |--Ocalemia P66b
    |--Strangalia maculata P66b, Im92
    |--Euryptera P66b
    |--Centrodera P66b
    |--Asilaris Pascoe 1866 P66b
    |    `--*A. zonatus Pascoe 1866 P66b
    |--Ephies Pascoe 1866 P66b
    |    `--*E. cruentus Pascoe 1866 P66b
    |--Desmocerus californicus B14
    |    |--D. c. californicus B14
    |    `--D. c. dimorphus Fisher 1921 B14
    `--Philus P66b
         |--P. antennatus [incl. P. inconspicuus] P66b
         |--P. globulicollis P66b
         |--P. pallescens Bates 1866 B66
         `--P. rufescens Pascoe 1866 P66b
  Phytoeciinae P66a
  Mesosinae P66a
  Apomecyninae P66a

Nomen nudum: Monohammus argentatus Hope in Masters 1887 M87

Bethelium signiferum Newm. 1840 [incl. B. flavomaculatum, B. fuscomaculatum, B. personatum Erichs. 1847] M87

*Blepephaeus succinctor (Chevrolat 1852) [=Monohammus succinctor; incl. M. obfuscatus White 1858, M. sublineatus White 1858] P66a

Ceresium simplex P66b [=Stenochorus simplex P66b; incl. Diatomocephala maculaticollis P66b, Oemona philippensis P66b, Ceresium vile Newm. 1842 M87]

Coptocercus biguttatus Donov. 1805 [=Callirhoe biguttata; incl. Co. senio Newm. 1841, Co. sexmaculatus, Co. tessellatus] M87

Coptocercus rubripes Boisd. 1835 [incl. C. allapsus Newm. 1841, C. assimilis Hope 1843, C. roei Hope 1843] M87

Coptomma Newman 1840 [incl. Navomorpha White 1855, Naomorpha (l. c.), Tmesisternus Audinet-Serville 1834 (n. n.)] SW03

Coptomma lineatum (Fabricius 1775) [=Callidium lineatum, Navomorpha lineatum, N. lineata (l. c.); incl. N. philpotti Brookes 1926, Coptomma textorium Newman 1840, N. textorium, N. textoria (l. c.)] SW03

Coptomma sulcatum (Fabricius 1775) [=Callidium sulcatum, Navomorpha sulcatum, N. sulcata (l. c.); incl. Coptomma acutipenne White 1846, N. acutipennis, N. neglectum Broun 1880, N. neglecta (l. c.)] SW03

Coptomma variegatum (Fabricius 1775) [=Callidium variegatum, *Tmesisternus variegatus, T. variegatum (l. c.); incl. *Co. virgatum Newman 1840] SW03

Hathliodes melanocephalus (Hope 1841) [=Hathlia melanocephala; incl. Hathliodes gracilis Blanch. 1853] M87

Olenecamptus bilobus [=Saperda biloba; incl. Authades indianus, Olenecamptus serratus Chevrolat 1835] P66a

*Type species of generic name indicated

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Last updated: 1 May 2017.

1 comment:

  1. Usually called longhorn beetles, rather than longicorn beetles, though either can be used ...

    ReplyDelete

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