Achilidae

Catonia pumila, copyright Judy Gallagher.


Belongs within: Auchenorrhyncha.

The Achilidae are a group of often flattened planthoppers whose nymphs live under bark or in cavities in dead wood (Carver et al. 1991).

Characters (from Carver et al. 1991): Apical segment of rostrum distinctly longer than wide. Fore wings usually folded flat, partially overlapping at rest, occasionally tectiform; clavus rather short, ending a little beyond half-way along fore wing, making a distinct angle with it; apex of confluent anal veins of fore wing reaching apex of clavus. Apex of second segment of hind tarsus with ventral row of small dark spines.

<==Achilidae
    |--Catanidia YS01
    |--Rhotala YS01
    |--Magadha Distant 1906 [Plectoderini] S04
    |--Achilus flammeus C91
    |--Aneipo CGW91
    |--Acixiites immodesta Hamilton 1990 RJ93
    |--Protepiptera P92
    |--Niryasaburnia Szwedo 2004 [Apatesonites] S04
    |    `--*N. burmitina (Cockerell 1917) [=Liburnia burmitina] S04
    |--Hooleya [Achillini] S04
    |    `--H. indecisa Cockerell 1922 S04
    |--Synecdoche S04
    |    |--S. dimidiata WW96
    |    `--S. impunctata WW96
    |--Epiptera WW96
    |    |--E. brittoni WW96
    |    `--E. variegata WW96
    `--Catonia WW96
         |--C. cinctifrons WW96
         |--C. picta WW96
         `--C. pumila WW96

*Type species of generic name indicated

REFERENCES

[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers vol. 1. Melbourne University Press: Carlton (Victoria).

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[S04] Szwedo, J. 2004. Niryasaburnia gen. nov. for ‘Liburnia’ burmitina Cockerell, 1917, from Cretaceous Myanmar (Burmese) amber (Hemiptera, Fulgoromorpha: Achilidae). Journal of Systematic Palaeontology 2 (2): 105–107.

[WW96] Wheeler, A. G., Jr & S. W. Wilson. 1996. Planthoppers of pitch pine and scrub oak in pine barrens communities (Homoptera: Fulgoroidea). Proceedings of the Entomological Society of Washington 98 (1): 100–108.

[YS01] Yoshizawa, K., & T. Saigusa. 2001. Phylogenetic analysis of paraneopteran orders (Insecta: Neoptera) based on forewing base structure with comments on monophyly of Auchenorrhycha (Hemiptera). Systematic Entomology 26: 1–13.

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