Brachystigmata

Male (above) and female Chlorogomphus magnificus, copyright Wildan Ardani.

Belongs within: Cavilabiata.
Contains: Paneurypalpidomorpha.

The Brachystigmata are a clade of dragonflies that mostly have fairly short pterostigmata in the wings (Bechly & Ueda 2002).

Synapomorphies (from Bechly & Ueda 2002): Wings with relatively short pterostigmata that cover only 1–3 complete cells (reversed in Libellulinae); “gaff” (= basal CuA before its branching) in hind wing strongly prolonged and very straight in the groundplan (sigmoidal in Eurypalpidomorpha); area between MP and CuA basally widened with more than one row of cells (reversed in some subclades); terminal branch of CuAa secondarily branched on CuA (inapplicable whe CuAa is secondarily unbranched); RP3/4 and MA strictly parallel up to the hind margin; area between RP2 and IR2 distally distinctly widened, with more than one cell row in the distal half (sometimes reversed).

<==Brachystigmata BU02
    |--Nannogomphidae BU02
    `--Paucipostnodalia BU02
         |--Prohemeroscopus Bechly et al. 1998 [Prohemeroscopidae] BU02
         |    |--*P. jurassicus BU02
         |    `--P. kuehnapfeli BU02
         `--Eubrachystigmata BU02
              |--Hemeroscopus R02 [Hemeroscopidae BU02]
              |    `--H. baissicus Pritykina 1977 R02
              `--Neobrachystigmata BU02
                   |--Paneurypalpidomorpha BU02
                   `--Chlorogomphida BU02
                        |--Araripechlorogomphus Bechly & Ueda 2002 [Araripechlorogomphidae] BU02
                        |    `--*A. muratai Bechly & Ueda 2002 BU02
                        `--Chlorogomphoidea BU02
                             |--Chloropetaliidae [Chloropetaliinae] BU02
                             `--Chlorogomphus SL89 [Chlorogomphinae BU02]
                                  |--C. brevistigma I92
                                  |    |--C. b. brevistigma I92
                                  |    `--C. b. okinawensis Ishida 1964 I92
                                  |--C. brunneus Oguma 1926 I92
                                  |    |--C. b. brunneus I92
                                  |    |--C. b. costalis I92
                                  |    `--C. b. keramensis Asahina 1972 I92
                                  |--C. hyalinus SL89
                                  |--C. iriomotensis Ishida 1972 I92
                                  |--C. magnificus SL89
                                  `--C. papilio RD77

*Type species of generic name indicated

REFERENCES

[BU02] Bechly, G., & K. Ueda. 2002. The first fossil record and first New World record for the dragonfly clade Chlorogomphida (Insecta: Odonata: Anisoptera: Araripechlorogomphidae n. fam.) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 328: 1–11.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[R02] Rasnitsyn, A. P. 2002. Special features of the study of fossil insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 8–12. Kluwer Academic Publishers: Dordrecht.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10^th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[SL89] Selys Longchamps, E. de. 1889. Odonates de Sumatra comprenant les espèces recueillies à Pulo Nias par M. le D.^r E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 444–484.