Amphientominae

Stimulopalpus japonicus, copyright Ashley Bradford.

Belongs within: Psocodea.

The Amphientominae are a group of scale-covered barklice in which the hind wing has vein M simple or is greatly reduced (New & Lienhard 2007).

<==Amphientominae [Amphientomini]
    |--Yinia Li 1995 L02
    |    `--*Y. capitinigra Li 1995 L02
    |--Yunientomia Li 2002 [=Yunamphientomia] L02
    |    `--*Y. ditaenia Li 2002 L02
    |--Lifashengia Lienhard 2003 T13 [=Obeliscus Li 2002 nec Beck 1837 nec Gray 1847 nec Popofsky 1913 L02]
    |    `--*L. zhongshani (Li 2002) [=*Obeliscus zhongshani] L03
    |--Stigmatopathus Enderlein 1903 NL07, T13
    |    `--*S. horvarthi Enderlein 1903 NL07
    |--Marcenendius Navás 1913 T13
    |    |--*M. nostras Navás 1913 L98
    |    `--M. illustris Navás 1923 L98
    |--Pseudoseopsis Badonnel 1955 L02
    |    |--*P. vilhenai Badonnel 1955 B55
    |    `--P. elegans New 1973 NL07
    |--Biocellientomia Li 2002 L02
    |    |--*B. bitrigata Li 2002 [=B. bitrigala] L02
    |    `--B. diplosticta Li 2002 L02
    |--Neuroseopsis Li 2002 L02
    |    |--*N. curtifurcis Li 2002 L02
    |    `--N. mecodichis Li 2002 L02
    |--Cornutientomus Li 2002 L02 [=Cornutientomius L03]
    |    |--*C. chinensis (Li 1993) [=Nephax chinensis, Seopsis chinensis] L02
    |    `--C. illepidotus Li 2002 L02
    |--Syllysis Hagen 1865 NL07, T13
    |    |--*S. (Syllysis) caudata [=Amphientomum caudatum] NL07
    |    |--S. samarangana Enderlein 1926 NL07
    |    `--S. (*Colposeopsis) sinipennis NL07
    |--Nephax Pearman 1935 T13
    |    |--*N. sofadanus Pearman 1935 T13
    |    |--N. angolensis Badonnel 1955 B55
    |    |--N. capensis Pearman 1935 T13
    |    `--N. fortunatus [=Perientomum fortunatum] L98
    |--Hemiseopsis Enderlein 1906 T13
    |    |--H. alettae Smithers 1989 T13
    |    |--H. fuellerborni S89
    |    |--H. machadoi Badonnel 1955 B55
    |    `--H. obscurus S89
    |--Amphientomum Pictet 1854 T13
    |    |  i. s.: A. ectostriolate Li 1999 L03, L02 [=A. ectostriolatis L03, A. ectostriolatum (l. c.) L03]
    |    |         A. elongatum M74
    |    |--*A. (Amphientomum) paradoxum Pictet 1854 L02
    |    `--A. (Palaeoseopsis Roesler 1944) B55
    |         `--A. (P.) flexuosum Badonnel 1955 B55
    |--Ancylentomus Li 2002 L02 [=Ancylentomius L03, Ancylopsocus Li 1997 non Tillyard 1926 L02]
    |    |--*A. fortuosus (Li 1997) [=*Ancylopsocus fortuosus] L02
    |    |--A. apicidealbatus Li 2002 L02
    |    |--A. ganquanensis Li 2002 L02
    |    |--A. longinervis Li 2002 L02
    |    `--A. macrurus (Li 1997) L03, L02 (see below for synonymy)
    |--Lithoseopsis Mockford 1993 [incl. Newseopsides Taylor 2013 (n. n.)] T13
    |    |--*L. hellmani (Mockford & Gurney 1956) T13 [=Pseudoseopsis hellmani M93]
    |    |--L. brasiliensis García Aldrete, da Silva Neto & Lopes Ferreira 2018 GASNLF18
    |    |--L. cervantesi García Aldrete 2004 T13
    |    |--L. chamelensis García Aldrete 2004 T13
    |    |--L. elongata (Mockford 1969) GASNLF18
    |    |--L. hystrix (Mockford 1974) T13 [=Amphientomum hystrix M93]
    |    |--L. humphreysi (New 1994) [=Seopsis humphreysi; incl. Newseopsides smithersi Taylor 2013 (n. n.)] T13
    |    |--L. incisa (Smithers 1989) [=Seopsis incisa] T13
    |    |--L. insularis García Aldrete 2004 T13
    |    `--L. tuitensis García Aldrete 2004 T13
    |--Paramphientomum Enderlein 1906 L02
    |    |--*P. (Paramphientomum) nietneri Enderlein 1906 NL07
    |    |--P. cordatum Li 2002 L02
    |    |--P. luzonica Banks 1937 N75
    |    |--P. malayense New 1971 NL07
    |    |--P. nigrceps Banks 1937 L02
    |    |--P. sinuosum New 1975 NL07
    |    |--P. striatum Thornton 1984 NL07
    |    |--P. triangulum Li 1992 L02
    |    |--P. (*Hormocoria) tristrigatum (Enderlein 1926) [=*Hormocoria tristrigata] NL07
    |    `--P. yumyum Enderlein 1907 L02
    |--Diamphipsocus Li 1997 L02
    |    |--*D. fulvus Li 1997 L02
    |    |--D. acaudatus Li 1997 L02
    |    |--D. concoloratus Li 2002 L02
    |    |--D. grammostictous Li 2002 L02
    |    |--D. magnimanubrus (Li 1995) L03, L02 [=Paramphientomum magnimanubrum L03, D. magnimanbrus (l. c.) L03]
    |    |--D. nanus (Li 1995) [=Paramphientomum nanum] L02
    |    |--D. shanxiensis Li 2002 L02
    |    |--D. signatus Li 2002 L02
    |    |--D. weihuai Li 2002 L02
    |    `--D. xanthocephalus Li 1997 L02
    |--Seopsis Enderlein 1906 L02
    |    |--*S. vasantasena Enderlein 1906 L02
    |    |--S. badia Li 2002 L02
    |    |--S. beijingensis Li 2002 L02
    |    |--S. concava Li 2002 L02
    |    |--S. cycloptera Li 1995 L02
    |    |--S. eucalla Li 2002 L02
    |    |--S. guibeiensis Li 2002 L02
    |    |--S. hirtella Li 1999 L02
    |    |--S. longisquama Li 2002 [=S. longisquamis] L03
    |    |--S. luzonica Banks 1937 NL07
    |    |--S. magna Li 2002 L02
    |    |--S. metallops S89
    |    |--S. metodicra Li 2002 L02
    |    |--S. multisquama Li 2002 [=S. multiquama] L03
    |    |--S. nanjingensis Li 2002 [=S. nanijingensis] L02
    |    |--S. nepalensis S89
    |    |--S. pavonia Badonnel 1955 S89, B55 [=S. pavonius B55]
    |    |--S. qinlingensis Li 2002 L02
    |    |--S. superba S89
    |    |--S. termitophila Badonnel 1955 S89, B55 [=S. termitophilus B55]
    |    `--S. tricolor Banks 1937 NL07
    `--Stimulopalpus Enderlein 1906 L02
         |--*S. japonicus Enderlein 1906 L02 [=Seopsis (Stimulopalpus) japonicus M93]
         |--S. acutipinnatus Li 2002 L02
         |--S. africanus Enderlein 1910 B55
         |--S. angustivalvus Li 2002 L02
         |--S. baeoivalvus Li 2002 L02
         |--S. biocellatus Badonnel 1949 B49
         |--S. brunneus (New 1975) NL07 [=Seopsis brunnea S95; incl. Se. harveyi Vaughan, Thornton & New 1989 S95]
         |--S. changjiangicus Li 1997 L02
         |--S. cochleatus Li 2002 L02
         |--S. concinnus Li 2002 L02
         |--S. conflexus Li 2002 L02
         |--S. distinctus Smithers 1995 NL07
         |--S. dolichogonus Li 2002 L02
         |--S. erromerus Li 2002 L02
         |--S. estipitatus Li 2002 L02
         |--S. exilis Li 2002 L02
         |--S. furcatus Li 2002 L02
         |--S. galactospilus Li 2002 L02
         |--S. heteroideus Li 2002 L02
         |--S. huashanensis Li 2002 L02
         |--S. immediatus Li 2002 L02
         |--S. introcurvus Li 2002 L02
         |--S. isoneurus Li 2002 L02
         |--S. medifascus Li 2002 L02
         |--S. mimeticus Li 1997 L02
         |--S. peltatus Li 2002 L02
         |--S. pentospilus Li 2002 L02
         |--S. phaeospilus Li 1999 L02
         |--S. polychaetus Li 2002 L02
         `--S. psednopetalus Li 2002 L02

Ancylentomus macrurus (Li 1997) L03, L02 [=Ancylopsocus macrourus L02, Ancylentomus macrourus (l. c.) L03, Ancylentomus maecrorurus (l. c.) L03]

*Type species of generic name indicated

REFERENCES

[B49] Badonnel, A. 1949. Psocoptères de la Cote d'Ivoire. Mission Paulian-Delamare (1945). Revue Française d'Entomologie 16: 20–46.

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[GASNLF18] García Aldrete, A. N., A. M. da Silva Neto & R. Lopes Ferreira. 2018. Lithoseopsis Mockford (Psocodea: Troctomorpha: Amphientomidae): a new species and first record for South America. Zootaxa 4526 (1): 91–95.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103–215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[N75] New, T. R. 1975. Lepidopsocidae and Amphientomidae (Psocoptera) from Malaysia and Singapore. Oriental Insects 9 (2): 177–194.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[S89] Smithers, C. N. 1989. Two new species of Amphientomidae (Insecta: Psocoptera), the first record of the family for Australia. Proceedings of the Linnean Society of New South Wales 111 (1): 31–35.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.

[T13] Taylor, C. K. 2013. The genus Lithoseopsis (Psocodea: Amphientomidae) in the Western Australian fauna, with description of the male of Lithoseopsis humphreysi from Barrow Island. Records of the Western Australian Museum Supplement 83: 245–252.