Thinoseius spinosus, copyright Steve Coulson.

Belongs within: Gamasina.
Contains: Pachylaelapidae, Leptolaelapidae, Macrochelidae, Parholaspidae.

The Eviphidoidea are a group of primarily soil- and litter-dwelling predatory mites.

Characters (from Lindquist et al. 2009): Adult dorsal shield entire; sternal shield of female with 3-4 pairs of setae, metasternal setae on small shields that are amalgamated with adjacent sternal-endopodal elements or present as discrete shields, occasionally inserted free in the integument; preendopodal platelets present or absent, female with a ventrianal or anal shield. Tibia I with 1 posteroventral seta, 2-3 posterodorsal setae, and 1 or 2 anterolateral setae (1(2) 3/2, 2(3)/1 2), tibia III with 7-8 setae (1(2) 1/1, 2/1 1). Palptarsal apotele 2-3-tined, epistome variously produced. Female podospermous, sacculus of the "A" or laelapid type. Male with genital aperture at anterior margin of a holoventral or sternitigenital shield; spematodactyl arises either from internal or external face of movable digit, free distally. Dorsal shield of deutonymph divided or entire; protonymph with 8-11 pairs of pygidial setae; larva with or without pygidial shield.

<==Eviphidoidea LKW09
    |  i. s.: Bulbogamasus [Bulbogamasidae] LKW09
    |           `--B. sinicus LKW09
    |--+--Pachylaelapidae K91
    |  `--+--Leptolaelapidae K91
    |     `--+--Macrochelidae K91
    |        `--Parholaspidae K91
    `--Eviphididae K91
         |  i. s.: Scamiphis LKW09
         |         Rafaphis LKW09
         |         Scarabacariphis grandisternalis LKW09
         |         Canestriniphis Potter & Johnston 1976 FH93
         |           `--C. megalodacne Potter & Johnston 1976 FH93
         |         Iphidiosoma S61
         |           |--I. bennwili Schweizer 1961 S61
         |           `--I. fimetarium S61
         |--Thinoseius Halbert 1920 [incl. Gammaridacarus Canaris 1962, Ligialaelaps Radford 1942; Thinoseiinae] ET79
         |    |--*T. berlesei Halbert 1920 ET79 [incl. Lasioseius fucicola Halbert 1920 ET79, T. fucicola S79]
         |    |--T. acuminatus ET79
         |    |--T. brevisternalis (Canaris 1962) [=Gammaridacarus brevisternalis] FH93
         |    |--T. hirschmanni P94
         |    |--T. kargi Hirschmann 1966 H66
         |    |--T. orchestoideae (Hall 1912) [=Seius orchestoideae] FH93
         |    |--T. schusteri Hirschmann 1966 H66
         |    `--T. spinosus (Willmann 1939) (see below for synonymy) FH93
         `--Eviphidinae KA90
              |--Scamaphis Karg 1976 ET79
              |    `--*S. equestris (Berl. 1911) [=Iphis (Pelethiphis) equestris; incl. S. exanimis Karg 1976] ET79
              |--Evimirus Karg 1963 ET79
              |    |--*E. uropodinus (Berlese 1904) ET79, FH93 [=Eviphis uropodina FH93]
              |    `--E. pulcherpori Karg 1989 FH93
              |--Crassicheles Karg 1963 ET79
              |    |--*C. concentricus (Oudemans 1904) [=Eviphis concentricus] ET79
              |    `--C. holsaticus ET79
              |--Scarabaspis Womersley 1956
              |    |--*S. inexpectatus (Oudemans 1905) ET79 [=Eumaeus inexpectatus ET79, Iphidozercon inexpectatus S61]
              |    |--S. orientalis (Berlese 1920) [=Iphidozercon orientalis; incl. S. aspera Womersley 1956] H98
              |    `--S. rykei LKW09
              |--Alliphis Halbert 1923 ET79
              |    |--*A. halleri (Canestrini & Canestrini 1881) ET79 (see below for synonymy)
              |    |--A. alpinus Schweizer 1961 S61
              |    `--A. oviforme (Schweizer 1949) [=Ameroseius oviforme] S61
              |--Copriphis Berlese 1910 ET79
              |    |--*C. pterophilus (Berl. 1882) [=Iphis pterophilus] ET79
              |    |--C. (Pelethiphis) dermanyssoides Berlese 1911 FH93
              |    |--C. elongatus Berlese 1911 FH93
              |    |--C. hastatellus RK91
              |    |--C. orbinellus Schweizer 1949 S61
              |    `--C. variola Schweizer 1949 S61
              `--Eviphis Berlese 1903 [incl. Iphidoides Oudemans 1904, Iphis Koch 1835] FH93
                   |--*E. pyrobolus (Koch 1839) [=Iphis pyrobolus, Eumaeus pyrobolus] ET79
                   |--E. drepanogaster LKW09
                   |--E. exemplaris Vitzthum 1925 V25
                   |--E. halleri S22
                   |--E. ostrinus (Koch 1835) [=*Iphis ostrinus] FH93
                   |--E. rufus Oudemans 1914 V25
                   `--E. stephaniniarus LKW09

*Alliphis halleri (Canestrini & Canestrini 1881) ET79 [=Gamasus halleri ET79; incl. A. alpinus Schweizer 1961 FH93, Copriphis (Pelethiphis) crinitus var. curtipilus Berlese 1911 FH93, Eviphis siculus Oudemans 1905 FH93, Alliphis siculus FH93]

Thinoseius spinosus (Willmann 1939) [=Lasioseius spinosus; incl. L. spinatus Sellnick 1940, L. uncinatus Sellnick 1940] FH93

*Type species of generic name indicated


[ET79] Evans, G. O., & W. M. Till. 1979. Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London 35: 139–270.

[FH93] Farrier, M. H., & M. K. Hennessey. 1993. Soil-inhabiting and free-living Mesostigmata (Acari-Parasitiformes) from North America: an annotated checklist with bibliography and index. North Carolina Agricultural Research Service, North Carolina State University, Technical Bulletin 302: i–xvi, 1–408.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[H66] Hirschmann, W. 1966. Gangsystematik der Parasitiformes. Teil 15. Gänge von Litoralmilben und neue Litoralmilbenarten. Acarologie: Schriftenreihe für Vergleichende Milbenkunde 9: 25–44.

[K91] Karg, W. 1991. The integration of morphologic, biogeographic and palaeogeographic studies to reveal the phylogeny of predatory mites. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 121–127. SPB Academic Publishing: The Hague.

[KA90] Krantz, G. W., & B. D. Ainscough. 1990. Acarina: Mesostigmata (Gamasida). In: Dindal, D. L. (ed.) Soil Biology Guide pp. 583–665. John Wiley & Sones: New York.

[LKW09] Lindquist, E. E., G. W. Krantz & D. E. Walter. 2009. Order Mesostigmata. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 124–232. Texas Tech University Press.

[P94] Pugh, P. J. A. 1994. Non-indigenous Acari of Antarctica and the sub-Antarctic islands. Zoological Journal of the Linnean Society 110: 207–217.

[RK91] Royce, L. A., & G. W. Krantz. 1991. A new rearing method for nematophagous mites. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 619–622. SPB Academic Publishing: The Hague.

[S79] Schuster, R. 1979. Soil mites in the marine environment. Recent Advances in Acarology 1: 593–602.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S61] Schweizer, J. 1961. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Parasitiformes Reuter, mit 246 Arten und Varietäten und 268 meist kombinierten Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84: i–vii, 1–207.

[V25] Vitzthum, H. 1925. Fauna sumatrensis. (Beitrag no. 5). Acarinae. Supplementa Entomologica 11: 1–79.

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