Pergamasus sp., copyright R. Penttinen.

Belongs within: Parasitidae.

Pergamasus is a genus of predatory mites found primarily in stable habitats such as forest and grassland soils. Females of this genus have the peritrematic shield fused with the holodorsal shield and both sexes have the opisthonotal region of the dorsal shield densely setose.

<==Pergamasus Berlese 1904 [incl. Plesiogamasus Hull 1918] S61
    |--P. (Pergamasus) FH93
    |    |--*P. (P.) crassipes (Linnaeus 1758) S61 (see below for synonymy)
    |    |--P. (P.) alpestris [=Gamasus (P.) alpestris] S22
    |    |--P. (P.) decipiens [=Gamasus (P.) decipiens] S22
    |    |--P. (P.) hamatus (Koch 1839) S22, D91 (see below for synonymy)
    |    |--P. (P.) lapponicus Trägårdh 1910 S22, D91 [=Gamasus (P.) lapponicus S22]
    |    |--P. (P.) longicornis (Berlese 1906) (see below for synonymy) FH93
    |    |--P. (P.) misellus Berlese 1903 S22, Koe91 [=Gamasus (P.) misellus S22]
    |    |--P. (P.) noster [=Gamasus (P.) noster] S22
    |    |--P. (P.) parvulus S22 (see below for synonymy)
    |    |--P. (P.) probsti [=Gamasus (P.) probsti] S22
    |    |--P. (P.) quisquiliarum (Canestrini & Canestrini 1882) S22, FH93 (see below for synonymy)
    |    |--P. (P.) runcatellus (Berlese 1903) (see below for synonymy) FH93
    |    |--P. (P.) runciger [=Gamasus (P.) runciger; incl. G. (P.) runciger var. armatus] S22
    |    `--P. (P.) theseus S22 [=Gamasus (P.) theseus S22; incl. P. theseus var. alpinus S61]
    `--P. (Thenargamasus) norvegicus (Berlese 1906) (see below for synonymy) FH93

Pergamasus incertae sedis:
  P. aequicornis Athias-Henriot 1967 FH93
  P. barbarus (Berlese 1905) [=Gamasus barbarus] H98
  P. basileus Schweizer 1961 S61
  P. brevicornis K09
  P. cornutus Schweizer 1961 S61
  P. crassicornutus D91
  P. cuneatus D91
  P. diessenhofenerus Schweizer 1961 S61
  P. digitulus Karg 1963 Koe91
  P. elegantulus Schweizer 1961 S61
  P. forazi Schweizer 1961 S61
  P. germanicus D91
  P. helveticus Schweizer 1961 S61
  P. homopodoides Athias-Henriot 1967 D91
  P. humusorum Schweizer 1961 S61
  P. lobatus Koh91
  P. mirabilis Willmann 1951 FH93 [=Amblygamasus mirabilis S61]
  P. neorunciger Schweizer 1961 S61
  P. obesus Koh91
  P. oxygynellus S61
  P. perlongum Schweizer 1961 S61
  P. petrophilus Schweizer 1961 S61
  P. pinetum Schweizer 1961 S61
  P. plenuspollex Schweizer 1961 S61
  P. serratulus Schweizer 1961 S61
  P. suecicus (Trägårdh 1936) D91
  P. truncus Schweizer 1961 S61
  P. trupchumi Schweizer 1961 S61
  P. valesianus S61
  P. vargasi Gorirossi 1955 FH93
  P. xerothermus Schweizer 1961 S61

Nomina nuda: Pergamasus arcuatus Dielmann 1991 D91
             Pergamasus carniolus Dielmann 1991 D91
             Pergamasus italicus Dielmann 1991 D91
             Pergamasus minutissimus Dielmann 1991 D91
             Pergamasus turriger Dielmann 1991 D91

*Pergamasus (Pergamasus) crassipes (Linnaeus 1758) S61 [=Acarus crassipes S61, Gamasus crassipes CF77, Siro crassipes G00; incl. Gamasus agilis CF77, Pergamasus crassipes var. alsacienne Dielmann 1991 (n. n.) D91, G. equestris CF77, G. fungorum CF77, G. quinquespinosus CF77, Acarus testudinarius Hermann 1804 CF77, G00, Gamasus testudinarius CF77, Siro testudinarius G00]

Pergamasus (Pergamasus) hamatus (Koch 1839) S22, D91 [=*Plesiogamasus hamatus S61; incl. Gamasus (Pergamasus) hamatus var. gracilis S22, Pergamasus hamatus var. longipes Schweizer 1961 S61]

Pergamasus (Pergamasus) longicornis (Berlese 1906) [=Gamasus (Pergamasus) crassipes var. longicornis; incl. G. (Pe.) magnicornutus Ewing 1909, G. (Parasitus) predator Banks 1910] FH93

Pergamasus (Pergamasus) parvulus S22 [=Gamasus (P.) parvulus S22; incl. P. parvulus var. denticulatus Schweizer 1961 S61, G. (P.) parvulus var. dilatellus S22]

Pergamasus (Pergamasus) quisquiliarum (Canestrini & Canestrini 1882) S22, FH93 [=Gamasus quisquiliarum FH93; incl. P. crassipes var. australicus Womersley 1942 H98]

Pergamasus (Pergamasus) runcatellus (Berlese 1903) [=Gamasus (P.) runcatellus; incl. P. neoruncatellus Schweizer 1961, P. teutonicus Willmann 1956] FH93

Pergamasus (Thenargamasus) norvegicus (Berlese 1906) [=Gamasus (Amblygamasus) septentrionalis var. norvegicus; incl. G. attenuipes Banks 1910, Hologamasus quinquedentatus Ewing 1920, Pergamasus septentrionalis Oudemans 1902] FH93

*Type species of generic name indicated


[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[D91] Dielmann, M. 1991. Computer in acarine taxonomy: a method for systematical analysis of the genus Pergamasus (Acari: Mesostigmata). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 239–248. SPB Academic Publishing: The Hague.

[FH93] Farrier, M. H., & M. K. Hennessey. 1993. Soil-inhabiting and free-living Mesostigmata (Acari-Parasitiformes) from North America: an annotated checklist with bibliography and index. North Carolina Agricultural Research Service, North Carolina State University, Technical Bulletin 302: i–xvi, 1–408.

[G00] Giribet, G. 2000. Catalogue of the Cyphophthalmi of the world (Arachnida, Opiliones). Revista Ibérica de Aracnología 2: 49–76.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[Koe91] Koehler, H. H. 1991. The impact of aldicarb on the succession of Gamasina; a one year study. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 749–758. SPB Academic Publishing: The Hague.

[Koh91] Kohn, M. 1991. Influence of the refuse dump biotopes on ecology of some gamasoid mites and ticks. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 199–204. SPB Academic Publishing: The Hague.

[K09] Krantz, G. W. 2009. Form and function. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 5–53. Texas Tech University Press.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S61] Schweizer, J. 1961. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Parasitiformes Reuter, mit 246 Arten und Varietäten und 268 meist kombinierten Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84: i–vii, 1–207.

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