Elassommatina

Oratemnus sp., copyright K. Edward.

Belongs within: Panctenata.
Contains: Chernetidae, Cheliferinae.

The Elassommatina is a clade of pseudoscorpions including species in which all legs have five post-coxal segments (with the tarsus undivided), and which have two or no eyes (Muchmore 1990).

<==Elassommatina MHG08
    |--+--Cheiridioidea MHG08
    |  |    |--Pseudochiridium M90 [Pseudochiridiidae MHG08, Pseudochiridiinae]
    |  |    |    `--P. insulae M90
    |  |    `--Cheiridiidae [Cheiridiinae] MHG08
    |  |         |--Cheiridium museorum M90
    |  |         `--Apocheiridium M90
    |  `--Sternophoridae [Sternophoroidea] MHG08
    |       |--Afrosternophorus MHG08
    |       |--Garyops MHG08
    |       |    |--G. depressus MHG08
    |       |    `--G. pumilus M90
    |       `--Idiogaryops M90
    |            |--I. paludis M90
    |            `--I. pumilis RS10
    `--Cheliferoidea [Atemnidae, Cheliferidae, Miratemnidae, Miratemninae] MHG08
         |  i. s.: Levichelifer [incl. Ocalachelifer] M90
         |           |--L. cribratus M90
         |           `--L. fulvopalpus M90
         |         Paratemnus elongatus M90
         |         Diplotemnus rothi M90
         |         Haplochelifer philipi M90
         |         Tyrannochelifer floridanus M90
         |         Ellingsenius sculpturatus M90
         |         Pugnochelifer amoenus M90
         |         Dactylochelifer M90
         |           |--D. copiosus M90
         |           `--D. silvestris M90
         |         Florichelifer aureus M90
         |         Parachelifer M90
         |         Xenochelifer davidi M90
         |         Aspurochelifer littlefieldi M90
         |         Hysterochelifer M90
         |         Phorochelifer mundus M90
         |         Idiochelifer nigripalpus M90
         |         Paisochelifer M90
         |           |--P. callus M90
         |           `--P. utahensis M90
         |--Chernetidae MHG08
         `--+--+--Caecatemnus MHG08
            |  `--Philomaoria [Philomaoriinae] MHG08
            |       `--P. pallipes (White 1849) [=Chelifer pallipes] T13
            `--+--+--+--Parazaona MHG08
               |  |  `--Withiidae MHG08
               |  |       |--Withius piger M90
               |  |       `--Juxtachelifer fructuosus M90
               |  `--Atemninae MHG08
               |       |--Cyclatemnus MHG08
               |       |--Oratemnus Beier 1932 MHG08, ABH14
               |       |    |--*O. articulosus (Simon 1899) ABH14
               |       |    |--O. afghanicus Beier 1959 ABH14
               |       |    |--O. boettcheri Beier 1932 ABH14
               |       |    |--O. brevidigitatus Beier 1940 ABH14
               |       |    |--O. confusus Murthy & Ananthakrishnan 1977 ABH14
               |       |    |--O. curtus (Beier 1954) ABH14
               |       |    |--O. distinctus (Beier 1948) ABH14
               |       |    |--O. loyolai Sivaraman 1980 ABH14
               |       |    |--O. manilanus Beier 1932 ABH14
               |       |    |--O. proximus Beier 1932 ABH14
               |       |    |--O. punctatus (Koch & Keyserling 1885) ABH14
               |       |    |--O. samoanus Beier 1932 ABH14
               |       |    |--O. semidivisus (Redikorzev 1938) ABH14
               |       |    |--O. timorensis Beier 1932 ABH14
               |       |    `--O. yodai Morikawa 1968 ABH14
               |       |--Stenatemnus Beier 1932 ABH14
               |       |    `--S. fuchsi (Tullgren 1907) [=Chelifer fuchsi] J98
               |       |--Titanatemnus Beier 1932 ABH14
               |       |--Anatemnus Beier 1932 ABH14
               |       |    |--*A. javanus (Thorell 1883) [=Chelifer javanus] ABH14
               |       |    |--A. cavernicola (Beier 1976) [=Oratemnus cavernicola] ABH14
               |       |    |--A. luzonicus Beier 1932 ABH14
               |       |    |--A. madecassus Beier 1932 ABH14
               |       |    |--A. nilgiricus Beier 1932 ABH14
               |       |    |--A. orites (Thorell 1889) ABH14
               |       |    |--A. subindicus (Ellingsen 1910) ABH14
               |       |    |--A. subvastus Alexander, Burger & Harvey 2014 ABH14
               |       |    |--A. subvermiformis (Redikorzev 1938) ABH14
               |       |    `--A. tonkinensis Beier 1943 ABH14
               |       |--Catatemnus Beier 1932 ABH14
               |       |    |--C. braunsi (Tullgren 1907) ABH14
               |       |    `--C. comorensis (Ellingsen 1910) ABH14
               |       |--Metatemnus Beier 1932 ABH14
               |       |--Tamenus Beier 1932 ABH14
               |       |--Mesatemnus Beier & Turk 1952 ABH14
               |       |    `--M. cyprianus ABH14
               |       `--Synatemnus Beier 1944 ABH14
               `--+--Miratemnus MHG08
                  `--Cheliferinae MHG08

*Type species of generic name indicated

REFERENCES

[ABH14] Alexander, J. B., M. A. A. Burger & M. S. Harvey. 2014. A new species of troglobitic Anatemnus (Pseudoscorpiones: Atemnidae) from the Pilbara bioregion of Australia. Records of the Western Australian Museum 29 (2): 141–148.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[M90] Muchmore, W. B. 1990. Pseudoscorpionida. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 503–527. John Wiley & Sones: New York.

[MHG08] Murienne, J., M. S. Harvey & G. Giribet. 2008. First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution 49: 170–184.

[RS10] Regier, J. C., J. W. Shultz, A. Zwick, A. Hussey, B. Ball, R. Wetzer, J. W. Martin & C. W. Cunningham. 2010. Arthropod relationships revealed by phylogenomic analysis of nuclear protein-coding sequences. Nature 463: 1079–1083.

[T13] Taylor, C. K. 2013. Further notes on New Zealand Enantiobuninae (Opiliones, Neopilionidae), with the description of a new genus and two new species. ZooKeys 263: 59–73.