Chioneini

Gonomyia sp., copyright John Rosenfeld.

Belongs within: Limnophilini.

The Chioneini are a group of crane flies characterised by bare eyes and legs lacking tibial spurs (Alexander & Byers 1981).

Characters (from Alexander & Byers 1981, as Eriopterini): Eyes glabrous. Terminal segment of palpus short; nasus absent. Tibial spurs absent. Wings, if present, with Sc₁ complete, free tip of Sc₂ absent, tip of R₁₊₂ present, R₄ and R₅ separate, CuA straight. Larva with plane of spiracular disc roughly perpendicular to long axis of body, disc surrounded by five lobes.

<==Chioneini [Chioneinae, Eriopterini] G09
    |--Neolimnophila AB81
    |--Cladura AB81
    |--Neocladura AB81
    |--Idiognophomyia AB81
    |--Tasiocera AB81
    |--Phantolabis lacustris AB81
    |--Gonomyodes AB81
    |--Gonomyopsis doaneiana AB81
    |--Gonempeda AB81
    |--Empedomorpha empedoides AB81
    |--Hesperoconopa melanderi AB81
    |--Arctoconopa AB81
    |--Hoplolabis armata WT11 [=Erioptera (Hoplolabis) armata AB81]
    |--Chionea AB81
    |--Aymaramayia G09
    |--Quechamyia G09
    |--Lipsothrix AB81
    |    |--L. neotropica G09
    |    `--L. nigrilinea AB81
    |--Eugnophomyia AB81
    |    |--E. apache G09
    |    `--E. luctuosa G09
    |--Cryptolabis (Cryptolabis) AB81
    |    |--C. (C.) magnistyla G09
    |    `--C. (C.) paradoxa G09
    |--Ormosia AB81
    |    |--O. (Ormosia) manicata AB81
    |    `--O. monticola AB81
    |--Cheilotrichia AB81
    |    |--C. (Cheilotrichia) alicia AB81
    |    `--C. (Empeda) G09
    |         |--C. (E.) stygia G09
    |         `--C. (E.) tanneri G09
    |--Teucholabis AB81
    |    |--T. (Teucholabis) complexa G09
    |    |--T. (Euparatropesa) G09
    |    |    |--T. (E.) invenusta G09
    |    |    `--T. (E.) lindneri G09
    |    `--T. (Paratropesa) collaris G09
    |--Toxorhina AB81
    |    |--T. (Toxorhina) G09
    |    |    |--T. (T.) magna AB81
    |    |    `--T. (T.) muliebris G09
    |    `--T. (Ceratocheilus) G09
    |         |--T. (C.) americana G09
    |         `--T. (C.) caledonica G09
    |--Molophilus AB81
    |    |  i. s.: M. fluviatilis Bangerter 1947 TKM93
    |    |         M. hirtipennis G09
    |    |--M. (Molophilus) G09
    |    |    |--M. (M.) gracilis G09
    |    |    `--M. (M.) plagiatus G09
    |    `--M. (Promolophilus) nitidus AB81
    |--Gnophomyia AB81
    |    |--G. jacobsoni G09
    |    |--G. lugubris G09
    |    |--G. rubicundula G09
    |    |--G. toschiae AB81
    |    |--G. triceps G09
    |    `--G. tristissima AB81
    |--Symplecta G09
    |    |--S. (Symplecta) G09
    |    |    |--S. (S.) cana G09 [=Erioptera (S.) cana AB81]
    |    |    |--S. (S.) hybrida G09
    |    |    |--S. (S.) macroptera G09
    |    |    `--S. (S.) stictica [=Erioptera (S.) stictica] AB81
    |    `--S. (Trimicra) pilipes G09 [=Erioptera (T.) pilipes AB81]
    |--Rhabdomastix AB81
    |    |  i. s.: R. neolurida AB81
    |    |         R. subfascigera AB81
    |    |--R. (Rhabdomastix) G09
    |    |    |--R. (R.) californiensis G09
    |    |    |--R. (R.) nuttingi AB81
    |    |    `--R. (R.) plaumanni G09
    |    `--R. (Sacandaga) AB81
    |         |--R. (S.) californiensis AB81
    |         `--R. (S.) monticola AB81
    |--Erioptera AB81
    |    |  i. s.: E. caloptera A16
    |    |         E. venusta A16
    |    |         E. vespertina A16
    |    |--E. (Erioptera) G09
    |    |    |--E. (E.) acucuspis G09
    |    |    |--E. (E.) cladophoroides G09
    |    |    `--E. (E.) septemtrionis AB81
    |    `--E. (Mesocyphona) G09
    |         |--E. (M.) costalis G09
    |         |--E. (M.) eiseni G09
    |         |--E. (M.) knabi G09
    |         |--E. (M.) melanderiana AB81
    |         `--E. (M.) troglodyta G09
    `--Gonomyia AB81
         |--G. (Gonomyia) G09
         |    |--G. (G.) kansensis G09
         |    `--G. (G.) subcinerea AB81
         |--G. (Leiponeura) G09
         |    |--G. (L.) haploa G09
         |    |--G. (L.) manca G09
         |    |--G. (L.) puer G09
         |    |--G. (L.) sulphurella G09 [=G. (Lipophleps) sulphurella AB81]
         |    `--G. (L.) sulphurelloides G09
         |--G. (Neolipophleps) G09
         |    |--G. (N.) alexanderi G09
         |    |--G. (N.) cinerea G09
         |    |--G. (N.) helophila G09
         |    `--G. (N.) strigilis G09
         `--G. (Paralipophleps) pleuralis AB81

*Type species of generic name indicated

REFERENCES

[A16] Alexander, C. P. 1916. New or little-known crane-flies from the United States and Canada: Tipulidae, Diptera, Part 2. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (3): 458–514, pls 16–21.

[AB81] Alexander, C. P., & G. W. Byers. 1981. Tipulidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 153–190. Research Branch, Agriculture Canada.

[G09] Gelhaus, J. K. 2009. Tipulidae (crane flies, tipúlidos). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 193–236. Ottawa, NRC Research Press.

[TKM93] Thaler, K., B. Knoflach & E. Meyer. 1993. Fragmenta faunistica Tirolensia—X. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 80: 311–325.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.