Cynipoidea

Ibalia leucospoides, from here.


Belongs within: Proctotrupomorpha.
Contains: Cynipidae.

The Cynipoidea is a clade of small wasps that appears to have been ancestrally parasitic on plant-boring insects (Grimaldi & Engel 2005).

Characters (from Grimaldi & Engel 2005): Metasoma laterally compressed; first and second metasomal sterna broadly fused; forewing median vein following distinctive course; forewing costa absent; antennal radicle absent.

<==Cynipoidea
    |  i. s.: Pseudeucoila bochei A71
    |         Paramblynotus MH11
    |         Paraspicera MH11
    |--Archaeocynipidae R02
    `--+--Austrocynipidae RN-A01
       `--+--Ibalia [Ibaliidae] RN-A01
          |    `--I. leucospoides A71
          `--+--Liopteridae RN-A01
             `--+--Eucoilidae A71
                |    |--Idiomorpha rapae [=Cothonaspis rapae] A71
                |    `--Eucoila [incl. Psilodora] A71
                |--+--Gerocynipidae GE05
                |  `--Cynipidae RN-A01
                `--Figitidae [Palaeocynipidae, Rasnicynipidae] RN-A01
                     |  i. s.: Aegilips A71
                     |--Parnips Ronquist & Nieves-Aldrey 2001 [Parnipinae] RN-A01
                     |    `--*P. nigripes (Barbotin 1964) [=Aulacidea nigripes] RN-A01
                     `--+--Thrasorinae RN-A01
                        |    |--Thrasorus RN-A01
                        |    |--Myrtopsen RN-A01
                        |    |--Plectocynips RN-A01
                        |    |--Pegacynips RN-A01
                        |    `--Euceroptres montanus RN-A01
                        `--+--Figitinae RN-A01
                           |    |--Melanips RN-A01
                           |    |--Figites A71
                           |    |--Lonchidia Thomson 1861 G30a, GM79
                           |    `--Thoreauella Girault 1930 G30a
                           |         `--*T. amatrix Girault 1930 G30a
                           `--+--Anacharitinae RN-A01
                              `--Charipinae [Allotriinae] RN-A01
                                   |--Charips victrix A71
                                   |--Hemicrisis RN-A01
                                   |--Phaenoglyphis japonicus (Ashmead 1904) (see below for synonymy) RN-A01
                                   |--Thoreauana Girault 1930 G30a
                                   |    `--*T. nativa Girault 1930 G30a
                                   |--Dilyta Foerster 1869 G30a, GM79
                                   |    `--D. mascagnini Girault 1935 G35
                                   `--Allotria Westwood 1833 GM79
                                        |--A. aphidae (Froggatt 1904) GM79 [=Hypodiranchis aphidae G30b]
                                        `--A. darci Girault 1933 G33

Phaenoglyphis japonicus (Ashmead 1904) [=Ceroptres japonicus, Euceroptres japonicus] RN-A01

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[G30a] Girault, A. A. 1930a. New pests from Australia. VIII. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 273-277).

[G30b] Girault, A. A. 1930b. New pests from Australia. IX. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 278-279).

[G33] Girault, A. A. 1933. Some beauties inhabitant not of commercial boudoirs but of Nature’s bosom, notably new insects. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 299-303).

[G35] Girault, A. A. 1935. Microhymenoptera Australiensis nova, mostly Chalcididae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 315-318).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

[R02] Rasnitsyn, A. P. 2002. Superorder Vespidea Laicharting, 1781. Order Hymenoptera Linné, 1758 (=Vespida Laicharting, 1781). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 242-254. Kluwer Academic Publishers: Dordrecht.

[RN-A01] Ronquist, F., & J. L. Nieves-Aldrey. 2001. A new subfamily of Figitidae (Hyenoptera, Cynipoidea). Zoological Journal of the Linnean Society 133: 483-494.

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