Belongs within: Apocrita.
Contains: Platygastroidea, Cynipoidea, Proctotrupidae, Chalcidoidea, Diapriidae.
The Proctotrupomorpha are a diverse clade of small wasps, most of which are parasitoids though several subgroups have become phytophagous.
See also: Taxon of the Week: To give Lovecraft nightmares.
Characters (from Grimaldi & Engel 2005): Lower tentorial bridge of head intervening between postgenae to reach occipital foramen; medial scutal sulcus absent.
<==Proctotrupomorpha
|--Mesoserphus GE05 [Mesoserphidae R02]
| `--M. karatavicus GE05
`--+--+--Stigmaphronidae GE05
| `--Ceraphronoidea MH11
| |--+--Lagynodes Forster 1841 MH11, GM79
| | `--+--Dendrocerus carpenteri MH11, M83
| | `--Megaspilus MH11 [Megaspilidae GE05]
| | `--M. luscipennis MH11
| `--Ceraphronidae [Calliceratidae] GE05
| |--Lygocerus A71
| |--Conostigmus A71
| |--Aphanogmus A71
| `--Ceraphron Jurine 1807 GM79
| |--C. achilles Dodd 1914 GM79
| |--C. bispinosus MH11
| `--C. javensis Girault 1917 G17
`--+--Platygastroidea MH11
`--+--Cynipoidea MH11
`--Proctotrupoidea MH11
|--+--Pelecinus [Pelecinidae] MH11
| | `--P. polyturator MH11
| `--Ropronia [Roproniidae] MH11
| `--R. garmani MH11
`--+--+--Heloridae R02
| | |--Protocyrtus R02
| | `--Helorus MH11
| `--+--Vanhornia [Vanhorniidae] MH11
| | `--V. eucnemidarum MH11
| `--Proctotrupidae MH11
`--Diaprioidea MH11
| i. s.: Jurapriidae R02
|--+--Monomachus [Monomachidae] MH11
| | `--M. australicus Girault 1925 G25
| `--+--Aneurhynchus MH11
| `--Coptera MH11
`--+--Chalcidoidea MH11
`--+--Maaminga Early, Masner et al. 2001 MH11, EM01 [Maamingidae]
| |--*M. rangi Early, Masner et al. 2001 EM01
| `--M. marrisi Early, Masner et al. 2001 EM01
`--+--+--Poecilopsilus MH11
| `--Entomacis MH11
`--+--Diapriidae MH11
`--+--Pantolytomyia ferruginea MH11
`--+--Ismarus MH11
`--+--Mymarommatidae [Mymarommatoidea] MH11
| |--Palaeomymar GE05 [incl. Petiolaria Bld. & Kryg. 1922 G30, GM79]
| | |--P. agapa R02
| | `--P. duisburgi GE05
| |--Mymaromma Girault 1920 MH11, GM79
| | `--M. anomalum MH11
| `--Mymaromella Girault 1931 MH11, G31
| |--*M. mira Girault 1931 G31
| `--M. mirissima Girault 1935 G35
`--Serphitidae [Serphitoidea] GE05
|--Aposerphites R02
`--Serphites R02
Proctotrupomorpha incertae sedis:
Peradeniidae EM01
Proctorenyxidae GE05
Neotelenomus G17
|--N. anthereae Dodd 1913 GM79
`--N. javae Girault 1917 G17
Eulagynodes Girault 1917 G17
`--*E. bicolor Girault 1917 G17
Odontacolus G17
|--O. australicus Dodd 1914 GM79
`--O. longispinosus Girault 1917 G17
Acoloides G17
|--A. arachnophagus Girault 1917 GM79
|--A. fasciatipennis Girault 1917 G17
|--A. javensis Girault 1917 G17
`--A. trispinosus Girault 1917 G17
Austroniidae R02
*Type species of generic name indicated
REFERENCES
[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.
[EM01] Early, J. W., L. Masner, I. D. Naumann & A. D. Austin. 2001. Maamingidae, a new family of proctotrupoid wasp (Insecta: Hymenoptera) from New Zealand. Invertebrate Taxonomy 15: 341-352.
[G17] Girault, A. A. 1917. New Javanese Hymenoptera. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 59-71).
[G25] Girault, A. A. 1925. Some gem-like or marvellous inhabitants of the woodlands heretofore unknown and by most never seen or dreamt of. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 191-193).
[G30] Girault, A. A. 1930. New pests from Australia. VIII. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 273-277).
[G31] Girault, A. A. 1931. A new habit in an old insect, Homo pudicus and new Eurytomidae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 280-283).
[G35] Girault, A. A. 1935. Microhymenoptera Australiensis nova, mostly Chalcididae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 315-318).
[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.
[R02] Rasnitsyn, A. P. 2002. Superorder Vespidea Laicharting, 1781. Order Hymenoptera Linné, 1758 (=Vespida Laicharting, 1781). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 242-254. Kluwer Academic Publishers: Dordrecht.
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